Trico tricolor tricolor (Hewitson, 1850), comb.n.

Euptychia tricolor Hewitson (1850, p. 440) . TL: river Amazon [Pará, Brazil]. Types: LT male (here designated): ‘ BMNH (E)-1,266,952//Type//B. M. TYPE No.

Rh. 3183 Euptychia tricolor ♂ Hew// Para. Hewitson Coll. 79 – 69. Euptychia Tricolor. Hew. 1’. (NHMUK) [figured on www.butterfliesofamerica.com, examined].

Trico tricolor tricolor

BRAZIL: Amapá: Serra do Navio (ICOMI), [0°59'N, 52°3'W], 14 Sep 1963, 1 ♀, (DZUP); Amazonas: 78 km W Manaus, Rio Preto de Eva, 100 m, ( Büche, M.), Dec 1996, 1 ♀ [MUSM-LEP-103501] , 1 ♀ [MUSM-LEP-103502], (MUSM); Camuna, (Le Moult, E.) , 1 ♁ [FLMNH-MGCL-266051], (FLMNH); Manaus, [3°7'S, 60°2'W], (Le Moult, E.), 1 ♁ [FLMNH-MGCL-266043], 1 ♁ [FLMNH-MGCL-266047], 1 ♁ [FLMNH-MGCL-266048], 1 ♁ [FLMNH- MGCL-266049], (FLMNH), ( Moss, A. M.) , 1 ♀ [BMNH (E)-525460], (NHMUK); Manaus, Reserva Ducke, [2°53'S, 59°57'W], ( Hutchings, R. W., Sullivan, J. B.), 13 Dec 1993, 1 ♁ [FLMNH-MGCL-266065], (FLMNH); Maués, [3°22'S, 57°43'W], Nov 1998, 1 ♁ [FLMNH-MGCL-266061], (FLMNH), (Balint), 1 ♁, (DZUP), (Le Moult, E.), 1 ♁ [FLMNH-MGCL-266044], 1 ♁ [FLMNH-MGCL-266045], 1 ♁ [FLMNH-MGCL-266050], 1 ♁ [FLMNH-MGCL-266062], 1 ♁ [FLMNH-MGCL-266063], 1 ♁ [FLMNH-MGCL-266064], (FLMNH); ' São Paulo de Olivença' - (error), (Moss, A. M.), 1 ♀ [BMNH (E)- 525449] , 1 ♀ [BMNH (E)- 525451], (NHMUK); Pará: [Rio] Tapajós, [4°16'8''S, 55°59'10''W], 25 m, 1 ♀ [BMNH (E)- 1668489], (NHMUK), (Bates, H. W.) , 1 ♁ [BMNH (E)- 1668486], (NHMUK), 3 Oct 2012, 1 ♁, (ZUEC), 4 Oct 2012, 1 ♁, (ZUEC), 8 Dec 2012, 1 ♁, (ZUEC); 15 km S Itaituba, [4°17'S, 56°5'W], (Callaghan, C. J.), 23 Jul 1978, 1 ♁ [FLMNH-MGCL-266054], (FLMNH); Amazonas, (Fassl, A. H.), (ZSM); Belém, Utinga, [1°27'S, 48°25'W], Feb 1957, 1 ♁, (DZUP) ; Rio Cumina, Igarapé dos Indios, 13 Oct 1936, 1 ♁, (DZUP) ; Itaituba on Rio Tapajós, [4°16'8''S, 55°59'10''W], (Le Moult, E.), 1 ♁ [FLMNHMGCL-266052], 1 ♁ [FLMNH-MGCL-266060], (FLMNH); Óbidos, [1°54'S, 55°31'W], (Le Moult, E.), 1 ♁ [FLMNH-MGCL-266053] , 1 ♁ FLMNHMGCL-266055], 1 ♁ [FLMNH-MGCL-266056], 1 ♀ [FLMNH-MGCL-266059], (FLMNH); Pará, [1°0'S, 51°11'W], 1 ♁ [BMNH (E)- 1266952], 1 ♁ [BMNH (E)- 1668482], 1 ♁ [BMNH (E)- 1668483], 1 ♁ [BMNH (E)- 1668485], (NHMUK), 15 Mar 1893, (ZSM), 1 ♀, (ZSM), (Bates, H. W.), 1 ♁ [BMNH (E)- 1668484], (NHMUK), ( Erhardt), (ZSM), (Mathan, M. de) , 1 ♁ [BMNH (E)- 1668480], 1 ♁ [BMNH (E)- 1668481], (NHMUK), (Moss, A. M.), 1 ♁ [BMNH (E)- 1499285], 1 ♁ [BMNH (E)- 525595], 1 ♁ [BMNH (E)- 525596], 1 ♁ [BMNH (E)- 525603], 1 ♁ [BMNH (E)- 525605], (NHMUK); Rio Tapajós, [4°16'8''S, 55°59' 10''W], 25 m, (Bates, H. W.), 1 ♁ [BMNH (E)- 1668487], (NHMUK), (Klug, G. G.), Sep 1931, 1 ♁ [BMNH (E)- 1499283], 1 ♁ [BMNH (E)- 1499284], (NHMUK); Santarém, [2°26'S, 54°43'W], (Le Moult, E.), 1 ♀ [FLMNH-MGCL-266057] , 1 ♀ [FLMNH-MGCL-266058], (FLMNH); Santa Bárbara do Pará, (Carneiro, E., Dolibaina, D., Dias, F., Moreira), 2-3 Feb 2010, 1 ♁, (DZUP), 6 Feb 2010, 1 ♁, (DZUP); Tapará, [2°2'S, 49°36'W], (Le Moult, E.), 1 ♁ [FLMNH-MGCL-266249], (FLMNH); Taperinha, [0°52' S,47°49'W], (Le Moult, E.) , 1 ♁ [FLMNH-MGCL-266046], (FLMNH) . FRENCH GUIANA: Cayenne: Approuage River, 10 mi W Regina, vicinity of St Anthanase Lodge, [4°11'10''N, 52°20'8''W], 120 m, (Brock, J.), 14-17 Oct 1990, 2 ♁, (BME) ; Cayenne: Cacao, [4°35' 51''N, 52°26'11''W], 0-150 m, (Harris, L. N.), 6 May 1998, 1 ♁ [FLMNH-MGCL-266088; Streams-falls-forest], (FLMNH) ; Cayenne, [4°56'N, 52°20'W], 1 ♁ [BMNH (E)- 1499279], 1 ♁ [BMNH (E)- 1499280], 1 ♁ [BMNH (E)- 1499281], 1 ♁ [BMNH (E)- 1668505], 1 ♁ [BMNH (E)- 1668507], 1 ♀ [BMNH (E)- 1668513], (NHMUK), (Bar, C.), 1 ♁ [BMNH (E)- 1668508], (NHMUK); St-Laurent du Maroni: Aloike-Maripasoula, [3°39'22''N, 53°59' 4''W], 110 m, (Brévignon, C.), 6 Apr 1987, 1 ♁ [MUSM-LEP-103500], (MUSM); Maroni River, (Le Moult, E.) , 1 ♁ [FLMNH-MGCL-266082; dissection, KW-14-015], 1 ♁ [FLMNH-MGCL-266083], 1 ♁ [FLMNH-MGCL-266084], 1 ♁ [FLMNH-MGCL-266085], 1 ♁ [FLMNH-MGCL-266086], 1 ♁ [FLMNH-MGCL-266087], 1 ♀ [FLMNH-MGCL-266089], 1 ♀ [FLMNH-MGCL-266090], (FLMNH); Maroni river, Maripasoula, [3°41'N, 54°2'W], (Brévignon, C.), 3 Mar 1987, 1 ♁ [MUSM-LEP-103499], (MUSM); Plateau Mines, Moutouchi Lodge, [5°19'47''N, 54°4'1''W], 45 m, (Sourakov, A.), 1-5 Mar 2013, 1 ♀ [FLMNH-MGCL-209668], (FLMNH); St. Laurent du Maroni, [5°30'N, 54°2'W], 1923, 1 ♁ [BMNH (E)- 1668510], (NHMUK), Jul-Sep 1915, 1 ♁ [BMNH (E)- 1668509], (NHMUK), Jun, 1 ♁ [BMNH (E)-1499282], (NHMUK), (Moult), Nov, 1 ♁ [BMNH (E)- 1668511], (NHMUK); Not located: ‘ French Guiana’, (Bar, C.), 1 ♁ [BMNH (E)- 1668506], (NHMUK) . GUYANA: Cuyuni-Mazaruni: Bartica, [6°24'N, 58°37'W], Mar 1939, 1 ♁, (BMB) ; junction of Cuyuni and Arimu rivers, 30 m, (Fratello, S.), 2 Oct 1991, 1 ♁ [FLMNH-MGCL-266076], (FLMNH) ; junction of Cuyuni and Arimu rivers, 46 m, (Fratello, S.), 28 Sep 1991, 1 ♀ [FLMNH-MGCL-266072], (FLMNH) ; East Berbice-Corentyne: Confluence of Oronoque and New River, [2°45'8''N, 57°26'5''W], (Hudson, C. A.), 20 Aug-20 Sep 1937, 1 ♁ [BMNH (E)- 1499271], (NHMUK) ; ' Kutari [River] Sources', [1°58'N, 56°36'W], (Hudson, C. A.), Jan-Feb 1936, 1 ♁ [BMNH (E)- 1668492], (NHMUK) ; Upper Demerara-Berbice: Essequibo River, 35 mi SW of Georgetown, Shanklands Resort, [6°28'44''N, 58°34' 54''W], 20 m, (Douglas, M. G.), 20-28 Sep 2006, 1 ♁ [FLMNH-MGCL-266074] , 1 ♁ [FLMNH-MGCL-266075], 1 ♀ [FLMNHMGCL-266073], 1 ♀ [FLMNH-MGCL-266078], 1 ♀ [FLMNH-MGCL-266080], 1 ♀ [FLMNH-MGCL-266081], (FLMNH); Not located: ‘ Guyana’, 1 ♁ [BMNH (E)- 1499272], 1 ♁ [BMNH (E)- 1668493], 1 ♁ [BMNH (E)- 1668495], (NHMUK); Demerara, [6°47'N, 58°10'W], (Castell), 1 ♁ [BMNH (E)- 1668491] , 1 ♁ [BMNH (E)- 1668494], (NHMUK); Essequibo River, Moraballi Creek, 10 Nov 1929, 1 ♁ [BMNH (E)- 1499270], (NHMUK) ; Potaro River, 1 ♀ [FLMNH-MGCL-266070], (FLMNH), (Roberts, C. B.), 12 Jul 1902, 1 ♁ [FLMNH-MGCL-266067], (FLMNH), 17 Jul 1902, 1 ♁ [FLMNH-MGCL-266068], 1 ♀ [FLMNH-MGCL-266069], (FLMNH), Jul 1902, 1 ♀ [FLMNH-MGCL-266079], (FLMNH), Oct-Nov 1902, 1 ♁ [FLMNH-MGCL-266066], (FLMNH); Río Demerara, 1 ♁ [BMNH (E)- 1499267], 1 ♁ [BMNH (E)- 1499268], 1 ♁ [BMNH (E)- 1499269], 1 ♁ [BMNH (E)- 1668490], (NHMUK); Suruwabaru CK, Mt. Wokamung, 680-690 m, (Fratello, S.), Nov, 1 ♀ [FLMNH-MGCL-266071], (FLMNH); Suruwabaru CK, Mt. Wokamung, 686 m, (Fratello, S.), Nov, 1 ♀ [FLMNH-MGCL-266077], (FLMNH) . SURINAME: Brokopondo: Berg-en-Dal, [5°9'N, 55°4'W], (Ellacombe, C. W.), May 1892, 1 ♁ [BMNH (E)- 1668496], (NHMUK), (Ellacombe, G. W.), May 1892, 1 ♁ [BMNH (E)- 1499273], 1 ♁ [BMNH (E)- 1499275], 1 ♁ [BMNH (E)- 1499277], (NHMUK); Marowijne: Río Lawa, Anapaika, [3°25'N, 54°1'W], (Malkin, B.), 27 Nov 1963, 1 ♁ [MUSM-LEP-103498], (MUSM); Not located: ‘ Surinam’, 1 ♁ [BMNH (E)- 1668497], 1 ♁ [BMNH (E)- 1668498], 1 ♁ [BMNH (E)- 1668499], 1 ♁ [BMNH (E)- 1668500], 1 ♁ [BMNH (E)- 1668502], 1 ♁ [BMNH (E)- 1668503], 1 ♁ [BMNH (E)- 1668504], 1 ♁ [BMNH (E)- 1668514], (NHMUK), (Ellacombe, G. W.), 1 ♁ [BMNH (E)- 1499276], (NHMUK); 'interior Surinam', Sep 1892, 1 ♁ [BMNH (E)- 1499278], (NHMUK), (Ellacombe, C. W.), Aug 1892, 1 ♁ [BMNH (E)- 1668501], (NHMUK), (Ellacombe, G. W.), Sep 1892, 1 ♁ [BMNH (E)- 1499274], (NHMUK) . TRINIDAD & TOBAGO: Not located: ‘Trinidad’, [10°26'17''N, 61°15'12''w], 1 ♁ [BMNH (E)- 1668512], (NHMUK) . VENEZUELA: Bolívar: El Dorado - Sta Elena, km 83, [6°9'59''N, 61°25' 37''W], 160 m, (Harris, L. & H.), 27 Jun 1984, 1 ♀ [FLMNH-MGCL-266042 ; km. 82], (FLMNH), (Lichy, R.), 18 Feb 1966, 1 ♁ [FLMNH-MGCL-266041; km. 82], (FLMNH) . Country unknown: Not located: ' Amazon', (Bates, H. W.), 1 ♁ [BMNH (E)- 1668488], (NHMUK) ; 'Amazonas', (Fassl, A. H.), 1 ♁ [BMNH (E)- 1499286], (NHMUK) ; no data, 1 ♁ [BMNH (E)- 1499287], 1 ♁ [BMNH (E)- 1499288], 1 ♁ [BMNH (E)- 1499289], 1 ♁ [BMNH (E)- 1499290], (NHMUK)

Other records: VENEZUELA: Bolívar: Río Grande, 90 km N El Callao, [7°56'54''N, 62°18'6''W], 100 m, (Neild, A. F. E.), 23 Oct 2000, 1 ♁, (ANNE) (Neild, A. (12 Aug 2020, pers. comm. to KRW by email with images)) .