Liogluta nitens (Mäklin in Mannerheim, 1852)

(Fig. 12 in Lohse & Smetana 1985; Figs. 98–100 in Klimaszewski & Winchester 2002)

Homalota nitens Mäklin in Mannerheim, 1852: 307.

Atheta (Philygra [sic!]) nitens: Bernhauer, 1907: 387 .

Atheta (Liogluta) insolens Casey, 1910a: 16, syn. nov.

Dimetrota (s. str.) resplendens Casey, 1910a: 104, syn. nov.

Atheta (s. str.) apposita Casey, 1911a: 90, syn. nov.

Atheta (Philhygra) nitens: Fenyes, 1920: 201 (as valid species).

Atheta (Liogluta) insolens: Fenyes, 1920: 209 (as valid species).

Atheta (Dimetrota) resplendens: Fenyes, 1920: 205 (as valid species).

Atheta (s. str.) apposita: Fenyes, 1920: 214 (as valid species).

Atheta (Philhygra) nitens: Bernhauer & Scheerpeltz, 1926: 629 (as valid species).

Atheta (Liogluta) insolens: Bernhauer & Scheerpeltz, 1926: 657 (as valid species).

Atheta (Dimetrota) resplendens: Bernhauer & Scheerpeltz, 1926: 665 (as valid species).

Atheta (s. str.) apposita: Bernhauer & Scheerpeltz, 1926: 638 (as valid species).

Atheta (Philhygra) nitens: Moore & Legner, 1975: 367 (as valid species).

Atheta (Liogluta) insolens: Moore & Legner, 1975: 364 (as valid species).

Atheta (Dimetrota) resplendens: Moore & Legner, 1975: 372 (as valid species).

Atheta (s. str.) apposita: Moore & Legner, 1975: 354 (as valid species).

Xenota nitens: Seevers, 1978: 670 (as valid species).

Liogluta insolens: Seevers, 1978: 263 (as valid species).

Dimetrota resplendens: Seevers, 1978: 260 (as valid species).

Xenota apposita: Seevers, 1978: 269 (as valid species).

Liogluta nitens: Lohse & Smetana, 1985: 288 (as valid species).

Atheta (Liogluta) nitens: Klimaszewski & Winchester, 2002: 40 (as valid species).

Type material. Lectotype of Homalota nitens (designated by Lohse and Smetana (1985)): UNITED STATES: Alaska: ♂, Sitka (Holmberg) (MZHF).

Lectotype of Atheta insolens (here designated): ♂, “QCI [ Queen Charlotte Islands]”, “ insolens ­2 PARATYPE USNM 39479” (red label), “CASEY bequest 1925” (NMNH) . Paralectotype: ♀, “QCI [ Queen Charlotte Islands]”, “ insolens Csy. ”, “TYPE USNM 39479” (red label), “CASEY bequest 1925” (NMNH) .

Holotype of Dimetrota resplendens: ♀, “QCI [ Queen Charlotte Islands]”, “ resplendens Csy. ”, “TYPE USNM 39123” (red label), “CASEY bequest 1925” (NMNH).

Lectotype of Atheta apposita (here designated): ♀, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “ apposita Csy. ”, “TYPE USNM 39308” (red label), “CASEY bequest 1925” (NMNH) . Paralectotype: ♀, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “ apposita PARATYPE USNM 39308” (red label), “CASEY bequest 1925” (NMNH) .

Additional material: UNITED STATES: Alaska: 7♀♀, Denali State Park, Byers Lake Campground, 23.vi.1978 (Smetana & Becker) ; 8♀♀, ditto but 25.vi.1978; ♀, ditto but above Byers Lake, 2500’, 24.vi.1978 (Smetana & Becker) ; 3♀♀, Denali State Park at Hwy. 1, 20.vi.1978 (Smetana & Becker) ; 4♂♂, 7♀♀, Kenai Mts., Ptarmigan Creek campground, 500–600’, 26.v.1978 (Smetana & Becker) ; 5♀♀, Kenai Mts., 16 mi. N Seward, 500–600’, 26.v.1978 (Smetana & Becker) ; 2♂♂, 15♀♀, Kenai Mts., 22 mi. N Seward, 600–800’, 27.v.1978 (Smetana & Becker) ; 7♀♀, Kenai Mts., 23 mi. N Seward, 600–900’, 28.v.1978 (Smetana & Becker) ; 10♀♀, Kenai Mts., 15 mi. N Seward, 400’, 29.v.1978 (Smetana & Becker) ; ♀, Kenai Mts., 2 mi. S Moose Pass, 500’, 30.v.1978 (A.Smetana & E.C.Becker) ; ♀, Kenai Peninsula, 1 mi. W Anchor Point, 1.vi.1978 (Smetana & Becker) ; ♂, 4♀♀, Kenai Peninsula, Anchor River Campground, 12 mi. N Homer, 450’, 3.vi.1978 (Smetana & Becker) ; 3♀♀, ditto but 5.vi.1978; 11♀♀, Kenai Peninsula, Anchor River at Highway 1, 450’, 4.vi.1978 (Smetana & Becker) ; 3♀♀, Kenai Mts., Crown Pt. Mine 4000’, 3 air mi. NE Crown Pt., 17.vi.1978 (Smetana & Becker) ; ♀, Kenai Mts., Tern Lake campground, 7000’, 12.vi.1978 (Smetana & Becker) ; ♀, Kenai Peninsula, Kasilof River at Hwy. 1, 12.vi.1978 (Smetana & Becker) ; ♂, ♀, Kenai Peninsula, 8 mi. SE Kasilof, 9.vi.1978 (Smetana & Becker) ; ♂, Kenai Peninsula, Kalifonsky Beach near Kenai, 8.vi.1978 (Smetana & Becker) ; 2♂♂, Kenai Peninsula, Clam Gulch, 6.vi.1978 (Smetana & Campbell) ; ♀, Alaska Range, Antimony Creek, 3500’, 2.5 mi. E mi. 181 George Parks Hwy., 27.vi.1978 (Smetana & Becker) ; ♀, 8 mi. NW Haines, 3.vii.1968 (Campbell & Smetana) (CNCI) ; Oregon: Benton Co .: 5♂♂, 2♀♀, Mary’s Peak, 3800’, 27.vii.1979 (J.M. & B.A.Campbell) (CNCI) ; Washington: ♀, Olympic N. P., Obstruction Peak, 6000’, 10.viii.1979 (J.M.Campbell & B.A.Campbell); ♂, ♀, Olympic N. P., 4.0–6.0 mi. SE Soleduck Campground, 2500–3100’, 15.viii.1979 (J.M.Campbell & B.A.Campbell) ; ♂, Olympic N. P., Lunch Lake, 4400’, 9 mi. SE Soleduck R.S., 17.viii.1979 (J.M.Campbell & B.A.Campbell) ; ♂, Olympic N. P., 7 mi. S Port Angeles, 2400’, 11.viii.1979 (J.M.Campbell & B.A. Campbell) (CNCI) ; CANADA: British Columbia: ♂, ♀, 5 mi. E Whistler’s Mt., 29.v.1968 (Campbell & Smetana) ; ♂, Slide Mt, 10 mi. E Barkerville, 4500’, 24.vi.1968 (Campbell & Smetana) (CNCI) .

Diagnosis. See Klimaszewski & Winchester (2002). This species is unusual in lacking a sclerotized spermatheca.

Discussion. The types of H. nitens, At. insolens, At. apposita and Dim. resplendens are similar in external characters, in the shape of the aedeagus (in male specimens) and in lacking a sclerotized spermatheca (in female specimens).

Distribution. Atheta nitens is known from Alaska, British Columbia, Washington and Oregon.