Rhinolophus lepidus Blyth, 1844

Rhinolophus lepidus Blyth, 1844: 486; Calcutta, INDIA (Collector unknown; Type unknown) [160] .

Rhinolophus refulgens Andersen, 1905: 124, pl. 4; Gunung Igari, Perak, MALAYSIA, 2000 ft. (A. L. Butlerm Esq., presenter; BM(NH) 1898.11.29.2) [161].

Rhinolophus lepidus refulgens [9].

Common English name: Blyth’s Horseshoe Bat

Barcode Index Number: BOLD:AAB9127 (5 DNA barcodes from Peninsular Malaysia; Fig 5)

Remarks: DNA barcodes of R. lepidus are associated with three BINs (BOLD:AAB9127, BOLD:ABZ1016 and BOLD:ABZ2266; Fig 5). The BIN, BOLD:ABZ1016 contains DNA barcodes recorded as R. lepidus and R. pusillus .

Some authors considered R. refulgens as a subspecies of R. lepidus [9, 11, 12] while some considered them to be distinct [50, 67]. Our NJ analysis (Fig 5) suggested that the DNA barcodes recorded as R. lepidus from Peninsular Malaysia may be distinct from DNA barcodes recorded as R. lepidus from Indochina. Similarly, Bumrungsri et al. [162] commented that R. lepidus from Peninsular Malaysia may represent a distinct taxon and the appropriate name would be R. refulgens based on the type locality. Due to the lack of DNA barcodes from the type locality of R. lepidus, India for comparison, we could not determine if DNA barcodes from Peninsular Malaysia represent the nominate R. lepidus or R. refulgens . Consequently, we tentatively retained only the name R. lepidus in this checklist (Fig 5).

IUCN status: Least Concern

Recorded at: Pahang: Krau Wildlife Reserve [11, 42], Kuala Atok, National Park [44], Lata Bujang Forest Reserve [56], Cameron Highland [60], Pulau Tioman [79], Bukit Ibam, Kemasul, Jengka, Klau Besar, Kenong and Gunung Aais [100]; Terengganu: Pasir Raja, Dungun [15], Tasik Kenyir [69], Gunung Tebu Forest Reserve [101]; Perak: Temengor Forest Reserve [46, 47, 111], Temenggor Lake [69]; Perlis: Wang Kelian State Park, [50], Gunung Stong State Park [100]; Selangor: Bukit Kutu Wildlife Reserve [51], Semangkok Forest Reserve [101]; Johor: Endau-Kota Tinggi Forest Reserve [56], Labis Forest Reserve [100]; Kedah: Ulu Muda Forest Reserve [57, 100], Bukit Hijau [100]; Kelantan: Air Panas-Gua Musang [61], Gunung Reng and Gua Musang [62]; Melaka: Unspecified [68]; Pulau Pinang: Bukit Panchor [100]; Negeri Sembilan: Gunung Angsi Forest Reserve [100, 101], Berembun Forest Reserve [101].

As R. refulgens: Perak: Maxwell Hill [23]; Pahang: Pulau Tioman [23], Krau Wildlife Reserve [41]; Johor: Pulau Pemanggil and Pulau Aur [23]; Negeri Sembilan: Pasoh Forest Reserve [45]; Kelantan: Gunung Stong State Park [67].

R. lepidus inhabits mature lowland and hill forests and has been reported roosting in caves and rock crevices, often with R. stheno [11, 14]