5. T. vernalis vernalis (Linnaeus, 1758)

Bercio & Folwaczny 1979 ( Geotrupes); Alekseev & Sakhnov 2000; Alekseev 2003 ( Geotrupes); Alekseev 2014.

oucher specimens: 3 [Dolzhanskoe, 27 June 1997; Pugachevo, 26 June 1997; Rybachii, 27 July 2008].

Kaliningrad Region: Gur. (Königsberg), Zel. (Rauschen); Bagr., Krasn., Nest., Zel. (Curonian Spit).

Comments: The sporadically distributed, occasional in the Kaliningrad Region species inhabited light pine forests and forest edges.