33. A. varians Duftschmid, 1805

[= niger (Panzer, 1797)]

Bercio & Folwaczny 1979 ( varians Dft. and niger Panz.).

Voucher specimens: 4 [23 km of the Curonian Spit, 30 May 2009; ibidem, 20 May 2012; Yantarny, 02 June 2014; Cherniakhovsk, 25 May 2018].

Kaliningrad Region: Chern. (Insterburg), Gur. (Metgethen, Dammhof, Forst Fritzen), Gur. (Königsberg for „ varians Dft. “), Zel. (Loppöhnen, Neuhäuser, Pillau, Rauschen, Rossitten).

Comments: The species names A. niger (Panzer, 1797) and A. niger Illiger, 1798 are considered homonyms and the specific name A. niger Illiger, 1798 for a widely distributed and endangered species of European dung beetle is conserved by suppression of other name (Krell et al. 2003). Panzer’s actual species is considered to be conspecific with A. varians Duftschmid, 1805 (Tamutis et al. 2011). All data from catalogue of Bercio & Folwaczny (1979) is united under name A. varians . The species is considered to be sporadically occasional in the Region.