Diaglyptidea conformis (Gmelin, 1790)

(New SJN: Fusahige-mame-togari-himebachi)

(Figs 7A–E, 62D, 65B)

Ichneumon conformis Gmelin, 1790: 2720 .

Hemiteles secernendus Schmiedeknecht, 1897: 108 .

Description based on Japanese materials. Female (n=60). Body length 3.4–4.8 mm. Body covered with silver setae.

Head. Clypeus covered with long and very dense setae, its anterior margin concave and indistinctly visible, obtuse in lateral view (Figs 4D, 62D). Face granulate (Fig. 7B). MSL 1.05–1.25 times as long as BWM. Malar space smooth with a few setae. Frons, vertex, and gena polished, with fine punctures. OOL almost as long as POL. Occipital carina incomplete dorsally, its lower end joined with hypostomal carina near mandibular base. Upper tooth of mandible almost as long as lower tooth. Base of mandible flat. Antenna with 21–24 flagellomeres, median and subapical parts not distinctly widened. FL III 3.2–3.75 times as long as maximum depth in lateral view.

Mesosoma. Lateral part of pronotum largely rugose except for dorsal and ventral smooth areas, with epomia (Fig. 7D). Mesoscutum matt (Fig. 7B) and subpolished, except for irregularly rugose posterior area, with distinct notaulus. Scutellum rugose-punctate anteriorly, punctate posteriorly. Mesopleuron covered with dense longitudinal striae except for smooth speculum (Fig. 7D). Metapleuron largely covered with sparse punctures and short striae, with a complete juxtacoxal carina. Anterior and posterior transverse carinae of propodeum complete (Fig. 65B). Area basalis distinct (Fig. 65B). Area superomedia distinct or partly indistinct laterally (Fig. 65B). Anterior part of propodeum finely and sparsely punctate. Median and posterior parts of propodeum covered with longitudinal, oblique and irregular rugae. Length of fore wing 2.9–4.2 mm. Areolet absent (Fig. 7E). Vein 2m-cu of fore wing with two bullae (Fig. 7E). Nervellus inclivous, intercepted behind the middle (Fig. 7E). Hind femur 4.0–4.5 times as long as maximum depth in lateral view. Hind TS I: II: III: IV: V = 2.0: 0.8: 0.6: 0.3: 0.5. Tarsal claws simple.

Metasoma. T I 1.4–1.5 times as long as maximum width, longitudinally striated. Median dorsal carina of T I present except for posterior part indistinct to absent. Dorsolateral carina of T I complete. T II punctate anterior 0.7 except for subapical transverse smooth area just in front of a shallow transverse concavity, anterior 0.4 covered with longitudinal striae. T III to T V covered with punctures except for smooth posterior areas. Ovipositor sheath 0.75–0.78 times as long as hind tibia. Ovipositor straight, with a nodus and ventral teeth.

Coloration (Figs 7A–E). Body (excluding wings and legs) black to blackish brown. Scape, pedicel, and base of FL I reddish brown to yellowish brown (but dorsal part sometimes darkened). Mandible yellowish brown except for darkened apex. Palpi, postero-dorsal corner of pronotum, and tegula yellow. Posterior and lateral margins of T II to T VII reddish brown. Membranous part of metasomal sternites and ovipositor reddish brown. Wings hyaline. Veins and pterostigma yellowish brown to brown. Legs reddish brown to yellowish brown. Trochanters, trochantelli, and fore and mid coxae yellow. Basal and apical parts of hind tibia and TS V more or less tinged with blackish brown. Dorsal part of hind coxa sometimes weakly tinged with dark brown.

Male (n=14). Similar to female. Body size overlapped with female but usually smaller than female. T I 1.6–1.7 times as long as maximum width.

Materials examined. JAPAN: KPM-NK 81641–81648, 6 F & 2 M, Tokyo, Izuoshima Is., Ohshima Town, Mt. Omaru, 17. VIII. – 5. X. 2012, K. Tsujii leg. ; KPM-NK 81598–81640, 32 F & 11 M, Tokyo, Miyakejima Is., Miyake Vil., Tsubota-rindo, 25. VIII. – 22. IX. 2012, K. Tsujii leg. (MsT) ; KPM-NK 81649, F, Tokyo, Hachijyojima Is., Mt. Miharayama, 31. V. 2012, K. Tsujii leg. ; KPM-NK 81650, F, Tokyo, Hachijyojima Is., Hachijo Town, Mitsune, Mt. Miharayama, 6. X. 2011, K. Tsujii leg. ; KPM-NK 81651–81653, 3 F, Tokyo, Mikurajima Is., Mikurajima Vil., Eigasawa, 20. IX. – 25. X. 2012, K. Tsujii leg. (MsT) ; KPM-NK 81654, F, Tokyo, Mikurajima Is., Mikurajima Vil., Borosawa, 13. V. 2012, K. Tsujii leg. ; KPM-NK 81655, F, Tokyo, Mikurajima Is., Mikurajima Vil., 14. V. 2012, K. Tsujii leg. (YPT) ; KPM-NK 81657, F, Nagano Pref., Outaki Vil., Mt. Ontakesan, Hakkaisan, 6. VIII. 2010, K. Watanabe leg. ; KPM-NK 81658, F, ditto, 7. VIII. 2010, K. Watanabe leg.; KPM-NK 81656, F, ditto, 30. VI. 2012, M. Ito leg.; KPM-NK 81659, F, Niigata Pref., Myoukou City, Suginosawa, Myoukou-sasagamine, 9. VII. 2013, S. Shimizu leg. ; KPM-NK 81660, M, ditto, 17. IX. 2013; KPM-NK 81662–81663, 2 F, Niigata Pref., Sado Is., Kanaishinbo, Hakuundai-Mt. Myoukenzan, 10. IX. 2010, K. Watanabe leg. ; KPM-NK 81664–81666, 3 F, Shizuoka Pref., Honkawane Town, Mt. Yamainudan, 14. VI. 2008, K. Watanabe leg. ; KPM-NK 81667–81668, 2 F, Toyama Pref., Nanto City, Togamura-kamimomose, 21– 28. VII. 2009 , M. Watanabe et al. leg. (MsT); KPM-NK 81669–81670, 2 F, Toyama Pref., Toyama City, Arimine, Jyuroudani, 7–14. VII. 2009 , M. Watanabe et al. leg. (MsT); KPM-NK 81671–81672, 2 F, ditto, 16–25. VIII. 2009; KPM-NK 81661, F, Osaka Pref., Takatsuki City, Mishimae, left bank of Yodo river, 27. X. 2012, S. Fujie leg. GERMANY: ZSM, F (det. Horstmann), Göttingen, 16. VIII. 1947 .

Distribution. Japan (Honshu, Sado Is., Izuoshima Is., Miyakejima Is., Hachijyojima Is., and Mikurajima Is.); widely distributed in Western Palearctic region.

Bionomics. Unknown in Japan. Some microlepidoptera and braconids have been recorded as the hosts (see Yu et al., 2016).

Remarks. This is the first record of this species from Japan and Eastern Palearctic region. This species can be distinguished from Diag. varipes by the occipital carina complete (incomplete medially in Diag. varipes) (Jonaitis, 1981).