Ceroderma asperata Raffray, 1890

(Figs. 1, 3–11, 19–21)

Ceroderma asperata Raffray, 1890b: 199 . Type locality: Penang Island, Malaysian Peninsula. Type material examined (MNHN). Holotype (Figs. 19–20): MALAYSIA: 3, labeled ‘Penang // MUSÉUM PARIS / 1917 / COLL. A. RAFFRAY // TYPE // C. asperata / A. Raffray det. // Ceroderma / asperata Raff / det. Löbl, 1970’.

Other material examined (NSMT). 2 3, 1 Ƥ, labeled ‘Ulu Gombak, Selangor / (termite nest) / [MALAY- SIA] / 27.vii.2003, M. Maruyama // Host: Coptotermes curvi - / gnathus Holmgren, 1913 / M. Maruyama det., 2003 // FAUNA MALAYSIA / Staphylinidae, Pselaphinae / Ceroderma asperata Raffray / S. Nomura det., 2003’.

Diagnosis. Head triangular; vertex largely concave; postocular margins roundly expanded laterally; triangular pronotum broadly incised at basal half; elytra evenly constricted at basal half, with three basal foveae; tergite IV slightly longer than V.

Redescription. Male. BL 2.66. Body (Figs. 1, 19) reddish brown, maxillary palpi and tarsi lighter; head, pronotum and elytra covered with dense setae, microtuberculate where setae arise. Head (Figs. 3, 19) much wider than long, HL 0.48, HW 0.81; vertex with tufted setae at anterior and posterior borders of large transverse median concavity, vertexal foveae and sulcus overlapped by concavity; postocular margins largely angularly expanded laterally, with dense long lateral setae; eyes well-developed, each composed of about 60 facets; antennomeres II–VIII (Fig. 4) of similar width, clubs formed by enlarged antennomeres IX–XI. Pronotum much wider than long, PL 0.55, PW 0.81; with shallow longitudinal median and lateral sulci; short discal carinae at basal half; lateral spines slightly larger than antebasal spines. Elytra wider than long, EL 0.82, EW 1.03; evenly constricted at basal half. Metaventrite concave along midline. Legs with femora thickened; protibiae (Fig. 5) angularly expanded mesally at apical half; mesotibiae (Fig. 5) with tiny membranous apical projection; metatrochanters (Fig. 7) with large elongate ventral projection. Abdomen wider than long, AL 0.81, AW 1.02; tergite IV with thick triangular lateral ridges; tergites V–VI with ridges much thinner; tergite VII (Fig. 8) with small lateral tubercles sharp. Sternite IV twice length of V at midline, with short median carina; sternites V–VII with short triangular basolateral ridges. Aedeagus (Figs. 9–11) well-sclerotized, length 0.37; broad basal bulb with large foramen, median lobe expanded apically.

Female. Similar to male in general. Measurements: BL 2.62, HL 0.47, HW 0.74, PL 0.55, PW 0.78, EL 0.78, EW 1.04, AL 0.82, AW 0.98. Vertex lacking setose concavity, widely separated foveae connected by shallow triangular sulcus in large impressed area; postocular margins less expanded than in male, lacking long lateral setae. Each eye composed of about 55 facets. Protibiae lacking mesal expansion at apical half; mesotibiae lacking apical tubercle.

Distribution. Malaysia: Penang, Selangor.

Host termite. Coptotermes curvignathus Holmgren (det. M. Maruyama).