Group niger

Definition. In imago and subimago of both sexes, tibiae of all legs blackish-brown, either entirely, or at least in proximal half (Figs 436–437, 467–469, 491–492, 526–527). This coloration is hypodermal and appears in larva of last instar prior to developing imaginal hypodermal coloration on femora.

Composition. Th. niger sp. n., Th. nigrabdominalis sp. n., Th. nigripes sp. n. and Th. nigrotibialis sp. n.