Forcipomyia (Metaforcipomyia) grandiseta n. sp.

(Figs. 46–51, 112, 141)

Diagnosis. Male: only species of Neotropical Forcipomyia (Metaforcipomyia) with a hind tarsal ratio of 2.17–2.20, flagellomere 10 1.10–1.35 X length of flagellomere 11, tarsomere 2 of the foreleg with thick, reinforced setae, and parameres present. Female: unknown.

Description of male. Head. Brown. Eyes abutting medially for length of 4–5 ommatidia. Antenna (Fig. 46) pale brown with plume setae well developed, flagellomeres 2–4 spherical, flagellomere 10 1.10–1.35 (1.20, n = 3) X longer than flagellomere 11, flagellomeres 10–13 elongate; flagellomere 13 with apical nipple, slightly constricted basally; AR 1.17–1.32 (1.25, n = 3). Palpus (Fig. 47) pale brown; segment 3 slightly swollen at midlength, with shallow sensory pit opening at midlength; PR 2.27–2.45 (2.37, n = 3). Labrum tapering apically.

Thorax. Uniformly dark brown; scutellum with approximately 8 strong setae and numerous minor ones. Legs brown, apex of hind tibia with 5 spines; tarsomeres 1–5 brown although progressively paler with pectinate scales; tarsomere 1 of foreleg with row of reinforced setae, tarsomere 2 of foreleg with 3–4 thick, basally setae (Fig. 48), tarsomere 3 of foreleg with one setae weaker that latter ones, tarsomere 1 of hind leg with two rows of spine-like bristles; foreleg TR 2.50–2.60 (2.55, n = 3), midleg TR 2.10–2.37 (2.23, n = 2), hind leg TR 2.17–2.20 (2.18, n = 3); claws curved, moderately stout. Wing (Figs. 49, 112) with abundant scales on costa, membrane infuscated with at most a dark patch on anterior margin of wing, distal to apex of radial cells; M2 poorly visible at base; first radial cell obliterated; second radial cell well developed; fork of cubitus situated at level of apex of costa; wing length 0.80–1.02 (0.90, n = 3) mm; breadth 0.26–0.32 (0.28, n = 3) mm; CR 0.44 (0.43–0.45, n = 3). Halter whitish.

Abdomen. Tergites brown. Genitalia (Fig. 50): Dark brown except gonostyli pale. Tergite 9 short, only extending to 1/4 length of gonocoxite; posterior margin nearly straight; cercus lobe-like, produced beyond midlength of gonocoxite; sternite 9 broad, with posterior margin with shallow posteromedian excavation. Gonocoxite stout, 1.80 X longer than greatest breadth; gonostylus 0.85 length of gonocoxite, almost straight, tip blunt. Parameres represented by a pointed posteromesally directed projection, gonocoxal apodemes slightly concave. Aedeagus (Fig. 51) stout, shield-shaped, lateral margins heavily sclerotized, posterior margin broad, with two apical lobes well developed, separated by narrow gap; basal arch low, heavily sclerotized; lateral arms nearly straight.

Female. Unknown.

Distribution. This species is known only from southern Costa Rica at 3–1320 meters (Fig. 141). The male from 3 km E of Cahuita was swept from a swamp forest just NW of the camping area in Cahuita National Park.

Taxonomic discussion. The reinforced setae on fore tarsomere 2 are shared with F. longiflagellata, and F. macroseta . Characters to distinguish all three species may be found in the key.

The aedeagus of this species is very similar to F. ronderosae, but in the latter species the apex of the aedeagus is concave.

Type material. Holotype male on microscope slide, labeled “ HOLOTYPE Forcipomyia (Metaforcipomyia) grandiseta Spinelli, Marino and Borkent, Costa Rica, Puntarenas prov., Golfito, Estación Agujas, 250–350 m, 15- VI / 15-VII-1999, J. Azofeifa, Malaise tp. LS276750 526550 #52838, CD 5156” (INBC). Paratypes, 2 males as follows: Costa Rica, 3 km E Cahuita, 29-X-1993, A. Borkent, CD 1641, 1 male (CNCI). Costa Rica, Puntarenas prov., Coto Brus, Progreso, Fca. Cafrosa, 2 km al NNE de la Escuela de Progreso, 1320 m, 23-XI-1995, M. Chinchilla, LS 318900N/595630E #7473, CD 5025, 1 male (MLPA).

Derivation of specific epithet. The name grandiseta – grand (large) seta refers to the stout setae on the foretarsomere 2 of the adult males of this species.

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