Forcipomyia (Metaforcipomyia) macroseta n. sp.

(Figs. 57–61, 113, 141)

Diagnosis. Male: only species of Neotropical Forcipomyia (Metaforcipomyia) with a hind tarsal ratio of 2.10, flagellomere 10 two X length of flagellomere 11, tarsomere 2 of foreleg with thick, reinforced setae, and parameres present. Female: unknown.

Description. Male. Head. Brown. Eyes abutting medially for length of four ommatidia. Antenna (Fig. 57) light brown except flagellomere 1 brown, with plume setae well developed, flagellomeres 2–4 elongate, flagellomere 10 two X longer than flagellomere 11, flagellomeres 10–13 elongate; flagellomere 13 with apical nipple, slightly constricted basally; AR 0.91. Palpus (Fig. 58) brown; segment 3 slightly swollen proximally (or on 2/3 proximal), with shallow sensory pit opening at midlength; PR 2.20. Labrum tapering apically.

Thorax. Uniformly dark brown; scutellum with 8 strong setae, and several minor ones. Legs brown, apex of hind tibia with 3–4 spines; tarsomeres with pectinate scales; tarsomere 1 of foreleg with row of dozen spine-like bristles, tarsomere 2 of foreleg with 1–2 thick reinforced setae, tarsomere 3 of foreleg with one setae weaker that latter ones, tarsomere 1 of hind leg with two rows of thick setae, tarsomeres 2–5 slightly paler; foreleg TR 2.75, midleg TR 2.00, hind leg TR 2.10; claws curved, moderately stout. Wing (Figs. 59, 113) with pattern of pigmented membrane, with dark patch on its anterior margin and at the apices of M2, CuA1 and CuA2; M2 only visible at wing margin; first radial cell obliterated; second radial cell well developed, triangular; fork of cubitus situated just at level of apex of costa; wing length 0.90 mm; breadth 0.30 mm; CR 0.42. Halter whitish.

Abdomen. Tergites brown. Genitalia (Fig. 60): Dark brown except gonostyli pale. Tergite 9 short, only extending to 1/4 length of gonocoxite; posterior margin broadly rounded; cercus lobe-like, produced beyond midlength of gonocoxite; sternite 9 broad, nearly straight. Gonocoxite stout, two X longer than greatest breadth; gonostylus 0.85 length of gonocoxite, almost straight, abruptly tapering to sharply pointed tip. Parameres represented by a slender, arrow-shaped, posteromesally directed projection, gonocoxal apodemes nearly straight, narrow. Aedeagus (Fig. 61) triangular, slightly tapering to pointed tip (triangular apex folded in the available specimen), arrowhead- shaped, sharply blunt tip; basal arch extending to 1/4 of total length; lateral arms extending laterally.

Female. Unknown.

Distribution. This species is known only from the type locality in Costa Rica, at 930 meters (Fig. 141).

Taxonomic discussion. This species shares with F. longiflagellata the presence of thick, reinforced setae on tarsomere 2 of the foreleg and the similarly shaped parameres. However, the wing of the latter species is plain and the aedeagus tapers slightly to a broad, truncate tip. It is also similar to F. grandiseta which shares the presence of thick, reinforced setae on tarsomere 2 of the foreleg, but differs from this species by the stouter aedeagus, which tapers to a broad posterior margin and bears two well developed apical lobes. The Nearctic species F. fehrerorum Grogan & Sigrist is nearly identical to F. macroseta, but the aedagus have an underlying hyaline pointed structure, the basal arch is lower and the apex is bifid.

Type material. Holotype male on microscope slide, labeled “ HOLOTYPE Forcipomyia (Metaforcipomyia) macroseta Spinelli, Marino and Borkent, Costa Rica, Limón prov., Talamanca, La Amistad NP entre Laguna Doroteri y Dabagri, 930 m, 23/ 27-VII-2007, A. Solis – M. Moraga, Malaise tp. LS 396680_543377 #92378” (INBC).

Derivation of specific epithet. The name macroseta – macro (large), seta refers to the stout setae on the foreleg tarsomere 2 of the male of this species.