Crassignatha pianma Miller, Griswold & Yin, sp. n.

urn:lsid:zoobank.org:act: AD950C98-9B09-40AC-85FE-04154475586F

Figs 67I, 72 C-D, 74-75, 76 A-B, 77, 78 A-D, 79 A-B, 98

Material Examined. Holotype: CHINA: Yunnan: Lushui Co., Pianma Township, Chang Yan He, 9.3 km ESE Pianma, 25.99363°N, 98.66651°E, 2470 m, 13-15 May 2005, mixed broadleaf deciduous and evergreen forest, dusting small webs near ground in forest understory, C. Griswold, CGY104 (CASENT 9022600, HNU), 1♁.

Paratypes: [same data as holotype] (CASENT 9029339, PV 0756-0758, CAS), 1♀; [same data as holotype] (CASENT 9029340, PV 0734-0735, CAS), 1 ♁; [same data as holotype] (CASENT 9029341, PV 0770-0778, CAS), 1♀; [same data as holotype] (CASENT 9022360, HNU), 2 ♁, 6 ♀, 1 juv; [same data as holotype] (CASENT 9022361, CAS), 3 ♁, 5 ♀, 1 juv.

Etymology. Named for the type locality.

Diagnosis. Male distinguished from other Crassignatha by the embolus shape (Fig. 75A), which is shorter than any other species.

Female distinguished by the procurved scape (Figs 76A, 79B) composed of two lobes of equal width (Figs 76B, 79A) in combination with ducts that do not run anteriorly between the spermathecae; other Crassignatha have the scape oriented ventrally to posteroventrally (Figs 76F, 79D) with the distal lobe of the scape more narrow than the basal lobe (Figs 76G, 79C) and/or with the ducts running anteriorly between the spermathecae before turning posteriorly (Figs 76I, 89B, D).

Description. Carapace orange-brown with small tubercles and sulci. Sternum orange-brown. Legs orange, femora I and II slightly swollen basally in female. Abdomen subspherical, light to medium gray, with numerous small sclerotized patches, some bearing long setae, male with single orange scutum laterally and posteriorly, female with small sclerite around spinnerets (Fig. 74).

Male palp: Median apophysis with two tapered distal processes. Embolus short, thick, and rigid, making a single turn (Fig. 75A, 77A).

Vulva: Scape slightly procurved (Figs 76A, 79B), with distal and basal lobes of equal width (Figs 76B, 79A). Spermathecae separated by their diameter. Ducts arise from posteromesal part of spermathecae, follow complex path mostly posterior to spermathecae before reaching openings near apex of scape.

Male (CASENT 9022360): Total length 0.86, carapace 0.43 long, 0.40 wide, clypeus 0.15, sternum 0.28 long, 0.26 wide, coxa IV separated by 2.15 times their width. Leg I: patella d1, tibia d2, metatarsus slightly sinuous, with two stiff bristles prolaterally; Leg II: patella d1, tibia d2, v3, metatarsus excavated proximoventrally; Leg III: patella d1, tibia d1; Leg IV: patella d1, tibia d1. Metatarsal trichobothria: TmI: 0.43; TmII: 0.44; TmIII: absent. Leg measurements: see Appendix A.

Female (CASENT 9022360): Total length 1.10, carapace 0.49 long, 0.45 wide, clypeus 0.11, sternum 0.30 long, 0.27 wide, coxa IV separated by 1.41 times their width. Leg I: patella d1, tibia d2; Leg II: patella d1, tibia d2; Leg III: patella d1, tibia d1; Leg IV: patella d1, tibia d1. Metatarsal trichobothria: TmI: 0.43; TmII: 0.46; TmIII: absent. Leg measurements: see Appendix A.

Natural History. This species builds a horizontal orb web (Fig. 72 C-D).