Scopaeus heronifer Frisch, spec. nov.

(Figs 14, 41, 86– 88, 132, 164)

Type specimens: Indonesia: Holotype ♂, Banten, Lebak: NE Majasari (06°37’32’’S, 106°23’53’’E), 460 m, 26.5.2016, leg. Frisch (MZB) . Paratypes (5 specimens): Jawa Barat: 1 ♀, Bogor, NO-slope Mt Salak (06°39’55’’S, 106°45’36’’E), 640 m, 13.9.2015, leg. Frisch (MZB) ; 1 ♀, Sukabumi, Cisalimas: Cisalimas River (E-slope Mt Halimun), (06°45’21’’S, 106°33’38’’E), 870 m, 19.9.2015, leg. Frisch (MZB) ; 1 ♂, Sukabumi, Sirnarasa, Cisarua: Ciawitali River (06°51’39’’S, 106°30’48’’E), 680 m, 20.9.2015, leg. Frisch (MFNB) ; 2 ♀, Sukabumi, Sirnarasa: Cimaja River (S-slope Mt Halimun), (06°51’32’’S, 106°31’06’’E), 670 m, 21.9.2015, leg. Frisch (MFNB, MZB) .

Description: Habitus and coloring as in Fig. 14. Head subquadrate. Penultimate antennal segment elongate. Mesotibia slender. Body surface with fine, dense, setose punctation, without microreticulation, shiny. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body medium brown with head and elytra somewhat darker than pronotum; antennae light brown with darker median segments; maxillary palpi and legs yellow-brown. Total body length 2.9 – 3.6 mm; forebody length 1.6 – 1.9 mm.

Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII with short, convex emargination in posterior seventh (Fig. 132). Aedeagus (Figs 41, 86 – 88) about 0.7 mm long with distal lobes asymmetrical and notably bent dextrad; dextral apical lobe slender in lateral view with distoventral incision (Fig. 86: arrow I) and tapered towards dorsad curved end (Figs 41, 86), in ventral and dorsal view convex laterally with retracted, narrow end (Figs 87, 88); sinistral apical lobe weakly sclerotized, evenly curved towards subacute apex (Figs 87, 88); deep distomedian emargination (Fig. 88: arrow III) dividing dorsal lobe in short, convex, dextral lobe and long, slender, dextrad curved, sinistral lobe angled bent towards truncate apex (Figs 87, 88); flagellum strongly projecting from apical lobes ventrally, evenly curved dorsad and projecting from apex of dextral apical lobe (Figs 41, 86), with strongly sclerotized (darker), in dorsal view acute-angled dextrad bent end, at point of bend with tooth pointing sinistrad (Fig. 87: arrow II); ventral lobe very short, triangularly projecting distad in lateral view (Figs 41, 86), with apical emargination in ventral view (Fig. 87); lateral lobes reduced, marked by lateroventral group of short setae (Figs 86, 87); median foramen elongate with acute proximal end, slightly shifted dextrad, with transverse ridge narrow and bent posteriad in middle (Fig. 87).

Female: Sperm pump stout with circular chamber (Fig. 164); bursa membranous.

Phylogeny: Scopaeus heronifer belongs to the S. longicornis complex (Frisch 2007: 204, 205) of the S. gracilis species group (Frisch et al. 2002: 39). This subgroup comprises two phylogeographical clades, the South African and Madagascan S. longicornis lineage and the Middle Eastern S. asirensis lineage (Frisch 2007: 213 – 215, 2015). Scopaeus heronifer, the first known Oriental species of the S. longicornis subgroup, represents another phylogenetic lineage. It lacks the characteristic, horn-shaped, sinistral extension of the base of the flagellum of the Afrotropical and Palaearctic species (Frisch 2007: e.g. Fig. 26). The sinistral tooth at the sharp apical bend of the flagellum (arrow in Fig. 87) is probably homologous to the sinistral enlargement of the flagellum of the S. asirensis lineage (Frisch 2015: e.g. Figs 9, 10).

Distribution: Scopaeus heronifer was collected at the edge of Mount Halimun-Salak National Park in the provinces of Banten and Jawa Barat.

Etymology: The epithet heronifer [adjective, Latin, composed of the noun heron (heron) and the verb ferre (to carry, to bring)] refers to the apex of the flagellum of the aedeagus (Fig. 94), which is reminescent of the head of a heron.