Scopaeus spiraliflagellatus Frisch, spec. nov.

(Figs 10, 35, 74 –76, 128, 157, 158)

Type specimens: Indonesia: Holotype ♂, Sulawesi Tengah, Poso: Tentena – Peura (Lake Poso), (01°47’59’’S, 120°38’08’’E), 550 m, 9.5.2017, leg. Frisch (MZB) . Paratypes (16 specimens): 1 ♀, same data as holotype (MFNB) . Sulawesi Tengah: 1 ♀, Poso: S Peura (Lake Poso), (01°51’48’’S, 120°38’59’’E), 520 m, 9.5.2017, leg. Frisch (MFNB) ; 1 ♀, Morowali: Betelene – Tinompo (02°06’06’’S, 121°08’26’’E), 430 m, 12.5.2017, leg. Frisch (MFNB) ; 1 ♂, 5 ♀, Poso: Tentena – Taripa (01°48’40’’S, 120°46’30’’E), 880 m, 10.5.2017, leg. Frisch (MFNB, MZB) ; 1 ♀, Poso: Tentena – Peura (Lake Poso), (01°47’59’’S, 120°38’08’’E), 550 m, 9.5.2017, leg. Frisch (MFNB) ; 1 ♂, 3 ♀, Poso: Palolo – Sedoa: NW Sedoa (01°18’02’’S, 120°16’22’’E), 1250 m, 6.5.2017, leg. Frisch (MFNB) . Sulawesi Selatan: 1 ♂, Makale, 17.07.1982, leg. Rougemont (HECO) . 1 ♂, labeled ‘ Célebès, Kandari 7’ (handwritten; Kendari in Sulawesi Tenggara?), ‘ apicipennis Fvl’ (handwritten; in litteris name) (ISNB) .

Description: Habitus and coloring as in Fig. 10. Total body length 3.1 – 4.0 mm; forebody length 1.9 – 2.2 mm. Head subcircular. Antennae with penultimate segment elongate. Mesotibia strongly thickened. Forebody surface with extremely spacious, fine, shallow, setose punctation, strongly shiny; abdomen also very finely, but more densely punctate than forebody, thus somewhat matter; body surface without microreticulation. Forebody setation comparatively long and strongly raised, including numerous, long macrosetae. Body black-brown except for yellow-brown posterior fifth of elytra, tip of abdomen, and legs with dark brown basal half of profemora becoming evenly lighter towards yellow-brown distal half; maxillary palpi darkened; antennae with scapus medium brown, pedicellus slightly darkened, segment 3 darkest, then evenly lighter towards light yellow-brown terminal segments.

Male:Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII widely emarginate in posterior fifth with median third of emargination almost semicircularly projecting posteriad (Fig. 128); macrosetae of sternite VIII long, conspicuous. Aedeagus (Figs 35, 74 – 76) about 1.2 mm long with distal lobes strongly asymmetrical; apical lobes remarkably enlarged apically with short, right-angled dorsad curved apices; dextral apical lobe ventroproximad extended, ending in dorsad pointing hook (Figs 35, 74); ends of apical lobes, in ventral and dorsal view, curved towards each other, together forming triangular end (Figs 75, 76); dorsal lobe subrectangular in lateral view (Fig. 74), with transverse, slightly oblique, truncate end in dorsal view (Fig. 76); flagellum extremely lengthened, whip-like, thin proximally, spiral with two windings at dextral side of aedeagus (Figs 35, 74 – 76); ventral lobe elongate, about four times as long as wide, pointing ventrodistad (Figs 35, 74); setose lateral lobes short, little projecting, strongly shifted distad to different extents (Figs 35, 74 – 76); median foramen large, asymmetrically extended distad with concave, distolateral enlargement at either side, without transverse ridge (Figs 74, 75).

Female: Sperm pump with apophysis adjacent to bursal duct; bursal duct strongly sclerotized in distal half, proximal half membranous (Figs 157, 158); bursa membranous.

Phylogeny: Scopaeus spiraliflagellatus is a member of the S. gracilis species group (Frisch et al. 2002: 39).

Distribution: Scopaeus spiraliflagellatus, probably widespread in Sulawesi, is recorded in the surroundings of Lake Poso, Sulawesi Tengah, and in Kendari, Sulawesi Tenggara.

Etymology: The epithet spiraliflagellatus [adjective, Latin, composed of the noun spiralis (spiral) and the adjective flagellatus (indicating a characteristic/feature of the flagellum)] refers to the spiral flagellum of the aedeagus of this new species (Figs 35, 74 – 76).