Scopaeus jacobsoni Cameron, 1930

(Figs 5, 30, 59 –61, 122, 148, 150)

Scopaeus jacobsoni Cameron, 1930: 346 .

Type specimen examined: Holotype ♂, Indonesia, Sumatra: Sumatera Barat, Bukittinggi; labeled ‘Holotype’ (red edged, round, white label), ‘ Fort de Kock (Sumatra) 920 M. 1925 leg. E. Jacobson. ’ (printed), ‘ Scopaeus jacobsoni Cam. TYPE’ (handwritten), ‘M. Cameron. Bequest. B.M. 1955-147.’ (printed), ‘Holotype / Scopaeus jacobsoni / Cameron 1930 / label by J. Frisch, 2022 ’ (subsequent holotype label, red, printed) (NHML) .

Cameron (1930: 346) described Scopaeus jacobsoni after a ‘unique’ specimen, which is a holotype by monotypy (ICZN, Article 73.1.2). The specimen labeled as holotype had already been labeled ‘TYPE’ by Cameron. Thus, a second specimen at NHML with the same locality label ( Fort de Kock, 1925, leg. E. Jacobson) subsequently labeled ‘Paratype’ is not a type specimen.

New Indonesian records: Bali: Tabanan: Pupuan Waterfall, 700 m, 14.6.1994, leg. Wunderle (PWCM) ; Tabanan: Batukaru, 17.6.1984, leg. Rougemont (HECO) . Banten: Lebak, E Bengkung: Cibeber (Cibareno River), (06°48’49’’S, 106°26’10’’E), 470 m, 22.9.2015, leg. Frisch ; Lebak: Citorek Kidul (06°44’51’’S, 106°19’13’’E), 870 m, 23.5.2016, leg. Frisch ; Lebak: Majasari (06°38’58’’S, 106°22’02’’E), 560 m, 25.5.2016, leg. Frisch ; Lebak: NE Majasari (06°37’32’’S, 106°23’53’’E), 460 m, 26.5.2016, leg. Frisch ; Lebak, Ciparasi: Ciparasi River (06°39’27’’S, 106°20’21’’E), 450 m, 28.5.2016, leg. Frisch. Jawa Barat: Bogor, NE-slope Mt Salak (06°39’55’’S, 106°45’36’’E), 640 m, 13.9.2015, leg. Frisch ; Bogor, Legok Dulang: Ciangsana River (06°43’08’’S, 106°31’15’’E), 1020 m, 18.9.2015, leg. Frisch ; Bogor, S Tenjolaja, Tapos: Ciampea River (06°41’36’’S, 106°42’24’’E), 1040 m, 2.10.2015, leg. Frisch ; Bogor, SW Pamijahan (Cimuara Herang River), (06°42’41’’S, 106°41’03’’E), 990 m, 3.10.2015, leg. Frisch ; Sukabumi, Cikaniki: Cikaniki River (Mt Halimun, Halimun-Salak NP), (06°44’46’’S, 106°32’25’’E), 1020 m, 17.9.2015, 5.10.2015, leg. Frisch ; Sukabumi, Cisalimas: Cisalimas River (E-slope Mt Halimun), (06°45’21’’S, 106°33’38’’E), 870 m, 19.9.2015, 14.5.2016, leg. Frisch ; Sukabumi, Sirnarasa: Cimaja River (S-slope Mt Halimun), (06°51’32’’S, 106°31’06’’E), 670 m, 21.9.2015, 15.5.2016, leg. Frisch ; Sukabumi, Sirnarasa, Cisarua: Ciawitali River (06°51’39’’S, 106°30’48’’E), 680 m, 20.9.2015, leg. Frisch ; Sukabumi, Simaresmi: Cisareno River (06°49’02’’S, 106°30’09’’E), 1000 m, 22.9.2015, leg. Frisch ; Sukabumi, Kiara Dua: Ciletuh River (07°08’40’’S, 106°36’55’’E), 670 m, 23.9.2015, leg. Frisch ; Sukabumi, Kiara Dua: Ciletuh River (07°08’27’’S, 106°37’46’’E), 710 m, 24.9.2015, 27.9.2015, leg. Frisch ; Sukabumi, N Djampang Kulon: Cikarang River (07°14’03’’S, 106°36’49’’E), 250 m, 25.9.2015, leg. Frisch ; Sukabumi, Simpenan: Cilulumpang River (07°08’43’’S, 106°38’24”), 740 m, 26.9.2015, leg. Frisch ; Sukabumi, NNW Cidahu: Cirasamala River (S-slope Mt Salak), (06°44’18’’S, 106°42’52’’E), 1210 m, 29.9.2015, leg. Frisch ; Sukabumi, NNW Cidahu: Javana Spa (S-slope Mt Salak), (06°44’43’’S, 106°42’51’’E), 1150 m, 20.9.2015, leg. Frisch ; Sukabumi, NE Pelabuhan Ratu (06°58’12’’S, 106°34’00’’E), 150 m, 19.5.2016, leg. Frisch. Jawa Timur: Pasuruan: Kakek Bodo Waterfall, 800 m, 20.9.1995, leg. Schillhammer (NHMW) . Sumatera Barat: Lake Toba, Samosir Island: Ambarita, 4.1977, leg. Jaccoud (MHNG) ; W Bukittinggi: Manindjau, 4.1977, leg. Jaccoud (MHNG) . Sumatera Utara: Simalungun: ‘ Prapat’ ( Parapat ?), 5.5.1984, leg. Rougemont (HECO) . Nusa Tenggara Barat: Lombok: Tetebatu, 6. – 10.8.1979, leg. Chambrier (MHNG) .

Redescription: Habitus and coloring as in Fig. 5. Head subquadrate. Antennae with penultimate segment quadrate. Mesotibia moderately thickened. Body surface with distinct, dense, setose punctation, without microreticulation. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Forebody reddish medium brown; abdomen darker brown with lighter brown tip; coloring of elytra variable, ranging from medium brown with indistinct, lateral darkening posterior of middle of length to contrasted color pattern with medium brown basis and gradually darker brown to blackish brown towards yellow-brown, posterior fifth to sixth of elytral length; maxillary palpi, antennae and legs yellow-brown. Total body length 2.6– 2.9 mm; forebody length 1.5– 1.6 mm.

Male:Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII with posterior fourth of posterior margin widely concave with slightly convex median fifth (Fig. 122). Aedeagus (Figs 30, 59 – 61) about 0.6 mm long; apical lobes in lateral view slender with acute apices, in ventral and dorsal view (Figs 60, 61) with right-angled, laterally projecting tooth at about middle of length, apically extended in slender lobes of unequal length somewhat curved mediad and widely separated by conspicuously transverse, apically truncate dorsal lobe extended in two long, ventrobasad pointing, diverging spines (Figs 59: arrow IV) with enlarged, overlapping ends; dextral spine of dorsal lobe evenly widened towards remarkably enlarged, transverse end with convex apical margin extended in huge, sinistrodistad curved tooth (Fig. 60: arrow V); sinistral spine of dorsal lobe less enlarged (Fig. 60: arrow VI); lateral lobes subrectangular, each with group of long, ventroapicad pointing and smaller group of shorter, ventrad pointing setae (Figs 59, 60); additional, unpaired, slender, apicad curved lobe projecting left of sinistral apical (lobe Fig. 60: arrow VII); ventral lobe moderately projecting, subtriangular with round apex (Fig. 59); median foramen wide, semicircular, with longitudinally broad, ventrally somewhat convexly projecting, transverse ridge (Fig. 30, 59, 60).

Female: Sperm pump with long, slender process segment; chamber short, circular, with short apophysis (Fig. 150); bursa mostly membranous with characteristic sclerotization at posterior end (Fig. 148).

Distribution: Scopaeus jacobsoni is widespread across Indonesia and recorded from Sumatra, Java and the Lesser Sunda Islands as far east as Lombok. I experienced the species to be very common in western Java.