Landouria abdidalem sp. nov.

urn:lsid:zoobank.org:act: B91C66B0-56B9-40EB-800E-4C321959C0B2

Figs 21, 35, 53, 56; Tables 3–4

Diagnosis

Landouria abdidalem sp. nov. is characterized by a sharply keeled shell, a long and narrow penis and epiphallus, and a crenated flagellum, which is shorter than the epiphallus and abruptly tapering at its proximal end.

Etymology

The name of the species is derived from the Javanese expression ʻAbdi Dalemʼ (used as a noun in apposition). Abdi Dalem is a person who devotes himself to serve the Keraton of Yogyakarta (Yogya Palace) and the Sultan.

Material examined

Holotype

INDONESIA • holotype (det. anat.); Yogyakarta, Kulonprogo, Jatimulyo, near Kiskendo Cave; 7°44′53″ S, 110°07′53″ E; 700 m a.s.l.; 25 Aug. 2017; B. Hausdorf leg.; D = 13.4 mm, H = 6.8 mm; MZB 21271.

Paratypes

INDONESIA – Central Java • 1 spec. (det. anat.); Purworejo, Donorejo, near Seplawan Cave, shrubs and limestone rock; 7°45′52″ S, 110°05′46″ E; 700 m a.s.l.; MZB 17733 • 3 spec.; same data as for preceding; MZB 17732 • 5 spec.; same data as for preceding; MZB 19314 • 4 spec.; same data as for preceding; ZMH 133410 • 1 spec.; same data as for preceding; ZMH 133421 • 10 spec.; Yogyakarta, Kulonprogo, Jatimulyo, near Kiskendo Cave; 7°44′53″ S, 110°07′53″ E; 700 m a.s.l.; ZMH 140026 • 1 spec. (det. anat.); Yogyakarta, Kulonprogo, Sabrang Kidul, snakefruit plantation; 7°44′38″ S, 110°07′55″ E; 730 m a.s.l.; MZB 17664 • 7 spec.; same data as for preceding; MZB 19253 .

Description

SHELL (Fig. 56; Tables 3–4). Depressed conical, with 5.75–6 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes sparse, mainly at periphery and at umbilical side; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus hardly eccentric, wide, comprising 33–48% of shell diameter, hardly obscured by columellar edge.

GENITALIA (Figs 21, 35; Table 4). Atrium short; penis long, becoming slightly broader proximally, without penis coecum; membranaceous sheath around distal part of penis connected with the epiphallus by tissue; penial retractor runs from diaphragm to proximal part of cylindrical epiphallus; flagellum shorter than epiphallus, broadest in its middle part, without node, abruptly tapering towards its blunt proximal end, crenated; vas deferens narrow and long; vagina long; oviduct shorter, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broader, spindle-shaped distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.

Remarks

With regard to the shell, L. abdidalem sp. nov. resembles L. tholiformis sp. nov. (Fig. 51), with which it clusters in the molecular phylogeny (Fig. 1). For differences, see remarks under that species. The range of L. abdidalem sp. nov. overlaps with that of L. zonifera sp. nov. For differences, see remarks under that species. Furthermore, the range of L. abdidalem sp. nov. overlaps with that of L. menorehensis sp. nov. (Fig. 88) from which it can easily be distinguished by the sharply keeled, depressed conical, brownishcorneous shell.

Distribution

Landouria abdidalem sp. nov. is endemic to the Menoreh Hills (Fig. 53).