Genus Cossulus Staudinger, 1887

Staudinger, 1887: 91.

Type species: Cossulus argentatus Staudinger, 1887 . Designation by monotypy.

Cossulus griseatellus Yakovlev, 2006 (Figs 21−22, 103) Yakovlev, 2006b: 14.

Type locality: NW Pakistan, Swat prov., Gabral-Tal.

Type material: holotype (male) in MNHW, examined.

Distribution: NW Pakistan, Swat prov., SO Afghanistan. Habitat and biology. Flight period: June-July. Altitude: 2100−2350 m. Material examined: male (paratype), SO Afghanistan, Safed Koh, Südseite, Kotkal, 2350 m, 21.06–1.07.1969, leg. Vartian (MNHW).

Cossulus herzi (Alpheraky, 1893) (Figs 23−24, 90, 104) Hypopta herzi Alpheraky, 1893: 346 .

Type locality: Samarkand [Uzbekistan, Samarkand].

Type material: lectotype (male) in ZISP, examined. Designation by Yakovlev (2006b).

Distribution: Kirgizstan, Uzbekistan, Tadzhikistan, N. Iran, Afghanistan (Daniel 1964b, 1965c, 1969b, 1971; Yakovlev 2006b).

Reported for Badakshan, Sarekanda; Anjuman Pass; Sarobi; Pandshir-Tal, Parian; Hazaradjat, Koh-i-Baba, Shah Fuladi; Pandshir-Tal, Chundsbu; Herat, Bala Murghab; Khurd-Kabul (Daniel 1964; 1969; 1971). Habitat and biology. Flight period: July −September. Altitude: 470−4200 m (Daniel 1964; 1969). Material examined: 27 males, Sarekanda, 4200 m, 31.0 7.1953, Badakshan, NO Afghanistan, Klapperich (ZSSM, MWM); 10 males, O. Afghanistan, Pandzhir Tal, Unteres Parian, 30.0 7.1961, Ebert (ZSSM); 1 female, Hazaradjat, Koh i Baba, Shah Fuladi, 3500 m, 3.0 7.1961, Ebert (ZSSM); 1 male, Khurd Kabul, 1900 m, 3.0 9.1965, Vartian (MWM); 2 males, Afghanistan, Bamian Prov., Band-e-Amir, 3100 m, 14.0 7.2013, leg. I. Pljustch, Yu. Skrylnik & O. Pak (RYB).

Cossulus intractatus (Staudinger, 1887) (Figs 25−26, 105)

Cossus intractatus Staudinger, 1887: 89 .

Type locality: Provinz Samarkand [Uzbekistan, Samarkand].

Type material (syntypes) in MHUB, examined.

Distribution: Kirgizstan, Uzbekistan, Tadzhikistan, Turkmenistan, Afghanistan (Yakovlev 2006b). Habitat and biology. Flight period: July. Altitude: 1000 m.

Material examined: 6 males, 1 female, N. Afghanistan, Prov. Badakhshan, Khwahan (Darwaz), 1000 m, 11.0 7.1972, leg. Brade & Naumann (MWM).

Cossulus kabulense (Daniel, 1965) (Figs 27−29, 88, 106)

Cossulinus kabulense Daniel, 1965: 142 .

Type locality: Afghanistan, Khurd-Kabul, 30 km SO v. Kabul.

Type material: holotype (male) in MNHW (coll. E. Vartian), examined. Distribution: Afghanistan (Paghman Mts.).

Habitat and biology. Flight period: June −July. Altitude: 2100−2200 m (Daniel 1965). Material examined: male (holotype) Afghanistan, Khurd-Kabul, 30 km SO v. Kabul, 2200 m, 20.06.–8.07.1963, Kasy & Vartian (MNHW); female (allotype), same data (MNHW); 2 males, Afghanistan, Pagman Gebirge, Kabul, 3000 m, Juni–Juli 1942, coll. Brandt (MNHS).

Cossulus putridus (Christoph, 1887) (Figs 30−32, 107)

Holcocerus putridus Christoph, 1887: 57 −58.

Type locality: Aschabad [Turkmenistan, Ashkhabad].

Type material (cotypes) in ZISP, examined.

Distribution: Turkmenistan, Iran, Afghanistan (Yakovlev 2006b).

Reported for Nuristan, Kutiau; Jalalabad (Daniel 1964; 1969).

Habitat and biology. Flight period: March −July. Altitude: 1550−3000 m (Daniel 1964). Material examined: 2 males, Salang pass, 3000 m, 7.0 7.72, Reshöft leg. (MWM); 1 male, 1 female, prov. Kadaghan, Salang pass, Nordseite, 11− 12.07.71, 2400 m, leg. Vartian (MNHW); 1 male, Badakhshan, Sarekanda, 21.0 7.1953, 2800 m, J. Klapperich leg. (MWM); 2 males, Salangpass, 3000 m, 9− 15.07.1973, leg. Flauger (MWM).

Cossulus strioliger (Alpheraky, 1893) (Figs 33−34, 89, 108)

Holcocerus strioliger Alpheraky, 1893: 346 .

Type locality: Samarkand [Uzbekistan, Samarkand].

Type material: lectotype (male) in ZISP, examined. Designation by Yakovlev (2006b). Distribution: Afghanistan, Iran, Tadzhikistan, Uzbekistan, Kirgizstan, Pakistan (Yakovlev 2006b). Reported for Polichomri (Daniel 1964), Paghman, 30 km WW Kabul (Daniel 1965) and Salang Pass (Daniel 1971). Habitat and biology. Flight period: May −July. Altitude: 700−2900 m (Daniel 1964; 1965). Material examined: male, Afghanistan, Pagman Gebirge, Kabul, 3000 m, Juni 1942, coll. Brandt (MNHS); 3 males, Salang pass, 3000 m, 7.0 7.72, Reshöft leg. (AHU; MWM); male, Bamyan, 12.0 6.1971, Dr. Reshöft leg.

(AHU); male, Kabul Fluss, Tang i Garu, Schlucht, 23.0 5.1977, leg. de Freina (MWM); male, 30 km NW of Kabul, 2500 m, Kasy & Vartian (MWM); 1 male, 1 female, Afghanistan, Bamian, Band-e-Amir, Sabzel vill., 2900 m, 19.0 7.2013, leg. I. Pljustch, Yu. Skrylnik & O. Pak (RYB); male, Kabul Fluss, Tang i Garu, 1700 m, 8.0 6.1972, Dr. Reshöft leg. (MWM).

Cossulus turcomanica (Christoph, 1893) (Figs 35−38, 109)

Hypopta turcomanica Christoph, 1893: 88 .

Type locality: Askhabad, Pul-i-Hatum, Tekke [Turkmenistan, Ashkhabad]. Type material: lectotype (male) in ZISP, examined. Designation by Yakovlev (2006b). Distribution: Turkmenistan, N. Afghanistan, NE Iran (Yakovlev 2006b).

The nominal subspecies was reported for Herat (Daniel 1964).

Material examined: male, Herat, 970 m, 5.5.56., Amsel (MWM); male, Salang pass, Kotale, leg. Dr. Reshöft, 3200 m, 14.0 7.72. (MWM); 24 males, female, Herat, 970 m, 5.0 5.1956, Amsel (ZSSM).

In eastern Afghanistan: subspecies albus (Daniel, 1971)

Cossulinus turkomanica albus Daniel, 1971: 658 .

Type locality: 40 km SW v. Kabul, Afghanistan.

Type material: holotype (male) in MNHW, examined.

Distribution: E. Afghanistan, N. Pakistan (Yakovlev 2006b).

Reported for Khurd-Kabul (Daniel 1971).

Habitat and biology. Flight period: April −May. Altitude: 970 m.

Material examined: 3 males, 1 female (holotype, paratypes) 17.0 6.1965, 40 km SW v. Kabul, 2300 m, Afghan., Kasy et Vartian (MWM); male, 35 km N. Kandaghar, Argandab Dan, 1150 m, 23.0 5.1961, Ebert (ZSSM); 2 males, 2 females, Pagman Gebirge, Kabul, 3000 m, Juni 1942, Brandt (MNHS); female, Khurd-Kabul, 2000 m, 11.0 6.71, leg. Vartian (MWM); 4 males, Tangi Gharu (Umg. Kabul) 14.0 6.72, leg. Dr. Reshöft (MWM); 7 males, Tangi- Schluht, 1800 m, 10.0 6.1975, leg. M. Dietz (MWM).

Cossulus habibae Yakovlev, Pljustch, Skrylnik & Pak, sp. nov. (Figs 39−41, 79, 89, 110) Type locality: Afghanistan, Bamian, Band-e-Amir, Sabzel vill. Type material: holotype (male) in ZISP.

Distribution: central Afghanistan.

Habitat and biology. Flight period: July. Altitude: 2800−2900 m.