Family * Calophyidae Vondráček, 1957

Comments

Burckhardt & Ouvrard (2012) admitted the artificial nature of their Calophyidae comprising five subfamilies. Two of these, Calophyinae and Mastigimatinae were included in the molecular analyses of Percy et al. (2018) and Cho et al. (2019), which both confirmed nonmonophyly of Calophyidae . The Mastigimatinae is removed here from Calophyidae and raised to family status. The other four subfamilies lack all metabasitarsal spurs. In addition, Atmetocraniinae Becker-Migdisova, 1973, Calophyinae and Metapsyllinae Kwon, 1983 bear an internal comb of apical metatibial spurs suggesting they may be closely related. Atmetocraniinae and Calophyinae share also the one-segmented asymmetric antennal flagellum in immatures (Burckhardt& Mifsud 2003; Burckhardt & Ouvrard 2012). With this, admittedly weak, evidence we keep the four subfamilies in the Calophyidae awaiting evidence to the contrary.