Digitonthophagus catta species group

Diagnosis. Pronotal punctures scattered and small on posterior half (Figs. 58, 93); surface behind anterior pronotal angles concave; male with two erected cephalic horns (Figs. 30, 32); females with anterior pronotal tubercles well developed and deeply obliquely concave (Figs. 94, 96); subapicodorsal lobe of FLP sclerite enlarged and sclerotized (Figs. 144, 147); FLP left surface with a conspicuous apophysis (Figs. 144, 147).