Key to studied Stachys taxa, based on nutlet morphological and anatomical characters

1. Nutlet color is brown, dark brown or dark brown-black ....................................................................................................................2

1. Nutlet color is taupe..........................................................................................................................................................................11

2. Nutlet color is brown ..........................................................................................................................................................................3

2. Nutlet color is dark brown or dark brown-black ................................................................................................................................9

3. Nutlet shape is ovatus, ovatus-rectangularis or ovatus-triangularis ...................................................................................................4

3. Nutlet shape is ellipticus............................................................................................................ S. megalodonta subsp. megalodonta

4. Ovatus-triangularis .............................................................................................................................................................................5

4. Ovatus, ovatus-rectangularis ..............................................................................................................................................................6

5. Nutlet surface ornamentation type is reticulate-foveate..................................................................................................... S. sivasica

5. Nutlet surface ornamentation type is verrucate ........................................................................................................... S. tundjeliensis

6. Nutlet surface ornamentation type is reticulate or reticulate-areolate................................................................................................7

6. Nutlet surface ornamentation type is verrucate or scalariform ..........................................................................................................8

7. Reticulate...................................................................................................................................................................... S. mardinensis

7. Reticulate-areolate........................................................................................................................................................... S. laetivirens

8. Scalariform ................................................................................................................................. S. megalodonta subsp. megalodonta

8. Verrucate...................................................................................................................................................................... S. baytopiorum

9. Nutlet shape is ellipticus-rectangularis............................................................................................................................... S. subnuda

9. Nutlet shape is ovatus or ovatus-rectangularis .................................................................................................................................10

10. Nutlet surface ornamentation type is alveolate ................................................................................................................. S. siirtensis

10. Nutlet surface ornamentation type is reticulate ................................................................................................ S. viscosa var. viscosa

11. Nutlet shape is ellipticus-rectangularis........................................................................................................................ S. benthamiana

11. Nutlet shape is ovatus-rectangularis........................................................................................................ S. viscosa var. odontophylla