326.

Formosan Tube-nosed Bat

Harpiola isodon

French: Murine isodonte / German: Goldgelbe Formosa-Rohrennase / Spanish: Ratonero isodonte

Other common names: Golden-haired Tube-nosed Bat, Taiwan Tube-nosed Bat

Taxonomy. Harpiola isodon Haochih et al., 2006,

“ Hualien County, Jhuosi Township, Yuli Wildlife Refuge [Taiwan], 23°32'N, 121°15'E,. 2,000 m asl.”

Limited genetic data place this species within Murina, close to M. harrisoni, M. hutton, and M. ussuriensis, making Murina paraphyletic and calling the generic validity of Harpiola into doubt. Monotypic.

Distribution. NW & C Vietnam (Hoang Lien National Park and Ngoc Linh National Reserve) and Taiwan.

Descriptive notes. Head-body 43-53 mm, tail 24- 5-36 mm, ear 12- 4-15 mm, hindfoot 5-:8-8- 1 mm, forearm 30-37- 3 mm; weight 3-4- 8 g. Fur is long and woolly; dorsally shiny golden yellow (hairs dark-brown-based with bright yellow bands and a dark brown tip, and dark brown guard hairs with shiny golden yellow tips mixed throughout); ventrally browner (hairs dark brown exceptfor light brown terminal two-thirds), sometimes with grayish white hairs along midline of chest and abdomen. Bright dorsal hairs continue onto bones ofwings, thumb, uropatagium, and feet. Last vertebra oftail extends beyond uropatagium. Wings and ears are dark brown, and face is furred except for long, tubular nostrils. Ears have distinct notch at upperthird of posterior border; tragus is moderately long but wide basally, gradually tapering to a backward-curved tip, with small, toothlike projection on outer margin. Skull is small, with narrow, shallow rostrum; braincase is elevated above rostrum with a distinctly higher mid-portion than the lambda; I* is bicuspid and enlarged; P* and P, are well developed, and are subequal to the canines and second premolars of the corresponding tooth row; C,is strongly bifid. Chromosomal complement has 2n = 44 and FN = 50 (Vietnam).

Habitat. Coniferous plantations or mixed forests of coniferous and broadleaved trees in Taiwan. It has been captured over streams in deciduous moss forests in Vietnam. Recorded at elevations of 1000-2400 m in Taiwan, and 1950-2250 m in Vietnam.

Food and Feeding. It has been seen foraging over stream backwaters and shallows flying c.0-2- 2 m above water or ground. It is a fast and very maneuverable flier.

Breeding. A pregnant female with one embryo was captured in May in Taiwan.

Activity patterns. No information.

Movements, Home range and Social organization. Two specimens were collected in a tunnel, where they may have been roosting.

Status and Conservation. Classified as Data Deficient on The IUCNRed List. Very little is currently known aboutits ecology. This species might be threatened by deforestation.

Bibliography. Kruskop (2013a), Kruskop & Shchinov (2010), Kruskop et al. (2006), Kuo Haochih et al. (2006), Nguyen Truong Son etal. (2015), Smith & Johnston (2008c).