307.

Small Woolly Bat

Kerivoula intermedia

French: Kérivoule de Bornéo / German: Kleine Wollfledermaus / Spanish: Querivoula de Borneo

Taxonomy. Kerivoula intermedia Hill & Francis, 1984,

“Lumerau, Sabah [Borneo, Malaysia], 5°12' N, 118°52' E.”

Relationship of K. intermedia to other species of Kerivoula is still uncertain; it might be closely related to K. minuta (likely given their identical karyotypes) or sister to the K. papillosa complex. Monotypic.

Distribution. Scattered localities in Peninsular Malaysia and N & SE Borneo; it might occur throughout the Thai-Malay Peninsula and the rest of Borneo.

Descriptive notes. Head-body c¢.34 36 mm, tail 33-42 mm, ear 9-14 mm, hindfoot 6-7 mm, forearm 26- 5-31 mm; weight 2:9—4- 2 g. Dorsal pelage is orange-brown; venter is paler. Face and lips are hairy, except nostrils. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavity just below tips on posterior borders; tragus is narrow and tall, with virtually straight anterior margin except for very slight convexity near tip, and has concave posterior margin with small hooked basal lobe. Membranes are light brown, with hairs scattered on posterior margin of uropatagium and no definite fringe; wings are attached at base of outer toes; and calcaris long. Skull is small and bulbous, averaging larger and less inflated than in the Least Woolly Bat, and posterior projection of palate can be longer than wide, with distal projection lacking in the Least Woolly Bat. Upper incisors are bicuspid; I, and I, are tricuspid, and I, is unicuspid; premolars are small and rounded; and talonid of M, is about the same width as trigonid. Chromosomal complement has 2n = 28 and FN = 52.

Habitat. Primary forest, rarely highly disturbed areas, and peat swamps (Borneo) from sea level up to elevations of ¢. 500 m.

Food and Feeding. No information.

Breeding. No information.

Activity patterns. Call shape is steep FM sweep, with start frequencies of 132-184-8 kHz (mean 173-1 kHz), end frequencies of 67-2-96-8 kHz (77-1 kHz), peak frequencies of 83-2-137-6 kHz (101-4 kHz), and durations of 0-7-3 milliseconds (1-7 milliseconds).

Movements, Home range and Social organization. No information.

Status and Conservation. Classified as Near Threatened on The IUCN Red List. Logging, agricultural expansion, and forest fires threaten the Small Woolly Bat throughoutits distribution. It is hard to capture and might be rare in much of its distribution; however, it is common in Krau Wildlife Reserve, Peninsular Malaysia.

Bibliography. Francis (2008a), Francis et al. (2007), Hill & Francis (1984), Husson & Morrogh-Bernard (2003), Hutson, Kingston & Francis (2008c), Khan et al. (2010), Kingston, Jones et al. (1999), Kingston, Lim & Zubaid (2006), Kumaran et al. (2006), Kuo Haochih et al. (2017), Yasuma & Abdullah (1997).