305.

Titania’s Woolly Bat

Kerivoula titania

French: Kérivoule titania / German: Titania-Wollfledermaus / Spanish: Querivoula titania

Taxonomy. Kerivoula titania Bates et al., 2007,

“Seima Biodiversity Conservation Area, Mondul Kiri Province, Cambodia, 12°10.49°N, 106°58.55'E, 290. m asl.”

Kerivoula titania appears to be sister to K. minuta, and this clade is sister to the rest of the hardwickii species group, although it might also be related to K. intermedia . Specimens previously attributed to K. fitania from Hainan and Taiwan islands now represent K. furva . Monotypic.

Distribution. E Myanmar, N Thailand, Laos, Vietnam, and E Cambodia.

Descriptive notes. Head—body 38- 4—41 mm, tail 45-8-53- 5 mm, ear 12- 5-15 mm, hindfoot 7-1-8- 3 mm, forearm 32-4-35- 9 mm; weight 4-7- 9 g. Dorsal pelage is brownish gray (hairs with black bases, light gray middles, and dark gray tips); venteris a little lighter (hairs black with whitish tips tinged with gray-brown). Face and lips are hairy, except nostrils. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavityjust below tips on posterior borders; tragusis narrow and tall, with virtually straight anterior margin except for very slight convexity near tip and concave posterior margin with small hooked basal lobe. Membranes are semi-translucent and gray, with hairs scattered on posterior margin of uropatagium and no definite fringe; wings are attached at base of outer toes. Fleshy lobe at base of thumb is oval and smooth in middle. Skull has broad, flattened braincase; rostrum has large narial pit and well developed rostral sulcus; and basioccipital pits are shallow. I? is unicuspid, with cingulum on its internal posterior border; I is equal in crown area to I?, with well-defined internally positioned cusp; canines are smooth, with deep longitudinal groove on outer sides; upper premolars are compressed in tooth row; M? lacks metastyle; I and I, are tricuspid, and I, is smaller with only well-defined central cusp and no lateral cusps; lower premolars are compressed in tooth row; and talonid of M,is reduced to ¢.70% of trigonid size.

Habitat. Lowland and montane primary and secondary evergreen and deciduous forests at elevations of 260-1600 m (Vietnam).

Food and Feeding. No information.

Breeding. Female Titania’s Woolly Bats in late pregnancy were caughtin early May in Ba Be National Park, northern Vietnam, and late April in Laos. Two pregnant females were captured in mid-May and two lactating females in early June in Laos, suggesting births occur in late May and possibly earlyJune.

Activity patterns. In Thailand, call shape is steep FM sweep, with start frequencies of 217-243 kHz (mean 227 kHz), end frequencies of 74-8-90 kHz (82-6 kHz), and peak frequencies of 137-7-168-7 kHz (157 kHz).

Movements, Home range and Social organization. No information.

Status and Conservation. Classified as Least Concern on The IUCN Red List. Titania's Woolly Bat does not seem to currently face any major threats, but becauseit is forest dependent, logging and agricultural development might be future threats.

Bibliography. Bates, Struebig et al. (2007), Chanson (2008), Douangboubpha et al. (2016), Kruskop (2013a), Kuo Haochih et al. (2017), Tu Vuong Tan et al. (2018), Wu Yi, Li Yuchun et al. (2012).