Enoggerinae Burks, new subfamily incertae sedis

Type genus.

Enoggera Girault, 1926.

Diagnosis.

Antenna with 9-12 flagellomeres, including either an incompletely divided clava or up to 4 clavomeres, sometimes including a small 4th clavomere. Temple absent, thus posterior margin of eye coincident with the posterior margin of the head dorsally (Fig. 65). Clypeus with transverse subapical groove. Labrum exposed, well-sclerotized, subrectangular, with marginal setae in a row. Mandibles with 3 teeth. Subforaminal bridge with postgena separated by lower tentorial bridge. Mesoscutellum with frenum indicated at least laterally, with axillular sulcus. Mesopleural area without an expanded acropleuron, with pits; mesepimeron not extending over anterior margin of metapleuron; two mesofurcal pits present. All legs with 5 tarsomeres; protibial spur stout and curved; basitarsal comb oblique. Metasoma with syntergum, therefore without epipygium. Egg parasitoids of Chrysomelidae ( Coleoptera).

Discussion.

Enoggerinae share a pair of mesofurcal pits with a number of other chalcidoid groups, including Herbertiidae, Moranilidae, and Asaphesinae . However, these groups are unstable in molecular analyses and do not form a clade (Cruaud et al., submitted), with Enoggerinae more often as the sister group of Coelocybidae . Therefore, Enoggerinae can be separated from all similar groups by the absence of the temple. With a different biology, Enoggerinae would also represent a discordant element if placed in any of the other groups.