Subgenus Apterodela Rivalier, 1950

Description. TL = 12.4–18.9 mm (mean = 16.16 mm, n = 95).

Labrum unidentate in both sexes (Figs 63–82, 85–87, 90–95, 98–100), rarely almost three-dentate in females (Figs 88, 89), or clearly tridentate in males (Figs 83, 84), with 3–11 long submarginal setae.

Pronotum indistinctly longitudinal or sub-quadrate (Figs 103–135, 138–140)—PW/PL = 0.81–1.09 (mean = 0.99, n = 204) (Fig. 269); wingless (Fig. 143).

Elytra with narrow and sloping shoulders, EW/EHW = 1.41–1.86 (mean = 1.57, n = 204) (Fig. 269); in one species with deep subapical sinuate notch (Figs 193–212), in the other species with poorly developed subapical sinuate notch (Figs 155–162, 165–179, 183–189) or entirely lacking (Figs 153, 154, 163, 164). White elytral pattern extremely reduced and presented by a small apical portion of humeral lunule (sub-humeral spot), a basal portion of apical lunule (supra-apical spot), and a short apical or centro-apical portion of middle band.

Internal sac of aedeagus with long cylindrical sharp mt (Figs 237–241, 251–255, 258–262, 265, 267, 271).