13. Lanonia magalonii (Henderson, Ninh Khac Ban & Nguyễn Quốc Dựng) Henderson & Bacon (2011: 893) .

Licuala magalonii Henderson, Ninh Khac Ban & Nguyễn Qu ốc Dựng (2008: 152). Type:— VIETNAM. Da Nang City, Hoa Vang District, Ba Na-Nui Chua Nature Reserve, 20 April 2007, A. Henderson, Nguyễn Qu ốc Dựng, Nguyen Canh, & Le Van Bo 3304 (holotype HN!, isotype NY!) .

Stems 0.7(0.3–1.5) m long, 2.3(2.0–2.5) cm diameter, solitary or clustered. Leaves 9(6–12) per stem; ligules 7.0 cm long, soon disintegrating into fibers; sheaths and proximalmost part of petioles with scattered, brown scales; petioles 38.7(33.0–43.0) cm long, 0.5(0.3–0.6) cm wide at the apex; petiole thorns usually poorly developed, brown or black, more or less regularly arranged on proximalmost part of petiole; hastulas flat, rounded, not infolded; leaf blades 46.0(44.0–48.0) cm wide; costas 10.2(6.3–16.2) cm long, narrow, with a pulvinus at the apex abaxially, with the few segments free except the middle pair joined at their bases; segments 4(3–6) per leaf, not mottled, with minute, reddish-brown scales abaxially, with (on dried specimens) straight margins; middle segments 31.3(23.5–38.5) cm long, 13.5(11.0–17.0) cm wide at the apex; apices of middle segments with adaxial splits not much deeper than abaxial ones. Inflorescences length not recorded; prophylls 18.2(16.2–22.0) cm long; peduncles 33.5(26.0–47.0) cm long; rachis bracts flattened, splitting apically and laterally, usually densely brown tomentose; rachillae not filiform nor thick and ribbed, sparsely to moderately covered with short, mostly simple, brownish hairs; staminate rachises absent or 11.5 cm long; staminate partial inflorescences 1–2, branched to 2-3 orders; staminate rachillae 4(3–5), 10.0(9.5–11.0) cm long, 1.4(1.1–1.6) mm diameter; staminate flowers 3.0 mm long; sepals not pedicellate at the base, not densely hairy; petals rounded at the apex; stamens with alternately long and short filaments and small, almost square anthers; filaments not elongate; pistillodes undivided; pistillate rachises absent; pistillate partial inflorescences 1, branched to 2–3 orders; pistillate rachillae on proximalmost partial inflorescence 6(2–9), 6.1(5.0–7.2) cm long, 1.4(1.1–1.7) mm diameter; pistillate flowers length not recorded, with short styles; fruits 7.3 mm long, 7.4 mm diameter, globose, color not recorded, with rough, pebbled surfaces; seeds with a curved basal intrusion.

Distribution and habitat:— Central Vietnam in Thua Thien-Hue province in Bac Ma National Park, and Da Nang City in Ba Na-Nui Chua Nature Reserve, in montane rainforest at 1,193(800-1,363) m elevation (Fig. 14).

Taxonomic notes:— Specimens determined as preliminary species Lanonia magalonii share a unique combination of qualitative variable states and are therefore recognized as a phylogenetic species.

Lanonia magalonii appears to be without obvious relatives. Hastulas are difficult to score; they are here scored as open and flat but some specimens appear almost infolded, perhaps by uneven pressing of the leaves.

Subspecific variation:— Lanonia magalonii occurs at higher elevations— 1,100 m on Ba Na-Nui Chua and 800– 1,363 m on Bach Ma—and the two mountains are separated from each other by a ca. 20 km lowland area. Specimens from the two mountains differ significantly from each other in two variables (petiole width, middle segment length) (t –test, P <0.05), with specimens from Ba Na–Nui Chua having lower values. Although there are too few specimens to test, those from Ba Na-Nui Chua have clustered stems 0.3–0.5 m long with 6-8 leaves per stem and 2–4 pistillate rachillae while those from Bach Ma have solitary stems 1–1.5 m long with 12 leaves per stem and 7–9 pistillate rachillae.

Sometimes plants in the field have been observed to have undivided middle segments.