Key to the Species of Hypatopa of Costa Rica

1. Male................................................................................................ 2

- Female............................................................................................. 50

2. Gnathos present)...................................................................................... 3

- Gnathos absent).......................................................................... Hypatopa rabio

3. Gnathos emarginate mesially (Figs. 139, 145, 149, 151, 153, 157, 159, 161, 163, 165, 167, 169, 171, 177, 179, 181, 183, 185, 189, 191, 195, 201, 203, 205, 209, 211, 213, 215, 219, 221, 223, 225, 231)........................................ 4

- Gnathos not emarginate mesially (Figs. 137, 147, 173, 141, 143, 155, 175, 187, 193, 197, 199, 207, 217, 227, 231, 233)... 36

4. Mesial emargination on a slightly raised or protuberant lobe (Figs. 139, 145, 153, 163, 171, 183, 189, 203, 205, 209, 221, 225, 231)................................................................................................ 5

- Mesial emargination on an entire margin (Figs. 149, 151, 157, 159, 161, 167, 169, 177, 179, 181, 185)................ 17

5. Apical process of ventral part of valva acutely curved in area from base to 1/2 (Figs. 139, 163, 209, 231)................ 6

- Apical process of ventral part of valva not acutely curved in area from base to 1/2 (Figs. 145, 153, 171, 183, 189, 203, 205, 221, 225)............................................................................................ 9

6. Apical process of ventral part of valva acutely curved laterally (Figs. 231)............................. Hypatopa rudis

- Apical process of ventral part of valva not acutely curved laterally (139, 163, 209).................................. 7

7. Proximal flange of valva longer than wide (Fig. 163); anellus rounded apically (Fig. 164)............... Hypatopa plebis

- Proximal flange of valva not longer than wide (Figs. 139, 209); anellus emarginate apically (Figs. 140, 210)............. 8

8. Gnathos deeply emarginate mesially; proximal flange wider than long (Fig. 139)...................... Hypatopa cladis

- Gnathos shallowly emarginate mesially; proximal flange about as wide as long (Fig. 209).................. Hypatopa io

9. Margin of proximal flange of valva crenulate (Figs. 183, 225)................................................. 10

- Margin of proximal flange of valva serrate (Figs. 145, 153, 171, 189, 203, 205, 221)............................... 11

10. Gnathos deeply emarginate mesially (Fig. 183); anellus truncate apically (Fig. 184)................... Hypatopa semela

- Gnathos shallowly emarginate mesially (Fig. 225); anellus deeply emarginate, forming two lateral lobes and with two porrect median spinelike processes (Fig. 226)......................................................... Hypatopa solea

11. Gnathos deeply emarginate mesially (Figs. 153, 171)........................................................ 12

- Gnathos shallowly emarginate mesially (Figs. 145, 189, 203, 205, 221).......................................... 13

12. Sclerite of phallus acutely curved near middle; vesica with barbed cornuti (Fig. 154)..................... Hypatopa crux

- Sclerite of phallus shallowly curved near middle; vesica without cornuti (Fig. 172).................... Hypatopa caepae

13. Phallus and sclerite of phallus longer than valva (Figs. 145, 204, 206)........................................... 14

- Phallus and sclerite of phallus shorter than valva (Figs. 190, 222)............................................... 16

14. Sclerite of phallus acutely sigmoid-shaped (Fig. 204)............................................... Hypatopa dux

- Sclerite of phallus not acutely curved throughout length (Figs. 146, 206)......................................... 15

15. Juxta divided mesially (Fig. 145); anellus nearly parallelsided throughout most of length (Fig. 146)........ Hypatopa cotis

- Juxta bandlike (Fig. 205); anellus constricted beyond base (Fig. 206)................................. Hypatopa erato

16. Gnathos widely emarginate (Fig. 189); anellus near parallelsided throughout most of length (Fig. 190)...... Hypatopa rego

- Gnathos narrowly emarginate (Fig. 221); anellus gradually narrowed from base (Fig. 222)............... Hypatopa umbra

17. Gnathos projected anteriorly near or greater than 1/2 width (Figs. 169, 177, 191, 195, 201, 219, 223).................. 18

- Gnathos not projected anteriorly as above (Figs. 149, 151, 157, 159, 161, 165, 167, 179, 181, 185, 211, 213, 215)........ 24

18. Ventral part of valva greatly protracted inwards basally (Figs. 169, 195, 23)...................................... 19

- Ventral part of valva slightly protracted inwards basally (Figs. 177, 191, 201, 19).................................. 21

19. Proximal flange of valva nearly twice as long as wide; apical process of ventral part of valva acutely curved basally (Fig. 169)....................................................................................... Hypatopa manus

- Proximal flange of valva about as long as wide; apical process of ventral part of valva not acutely curved from base (Figs. 195, 223)............................................................................................... 20

20. Apical margin of proximal flange wider than base (Fig. 223); anellus widely emarginate, forming two lateral lobes, and with two convergent apical processes (Fig. 224)...................................................... Hypatopa texo

- Apical margin of proximal flange narrower than base (Fig. 195); anellus rounded apically (Fig. 196)........ Hypatopa verax

21. Gnathos deeply emarginate mesially (Figs. 177, 191, 219).................................................... 22

- Gnathos shallowly emarginate mesially (Fig. 201).................................................. Hypatopa nox

22. Sclerite of phallus recurved apically (Figs. 192, 220)......................................................... 23

- Sclerite of phallus not recurved apically (Fig. 178).............................................. Hypatopa scobis

23. Margin of proximal flange serrate (Fig. 191); sclerite of phallus recurved to near base of anellus (Fig. 220).... Hypatopa ira

- Margin of proximal flange entire (Fig. 219); sclerite of phallus recurved to subapical region (Fig. 192)...... Hypatopa styga

24. Ventrolateral margin of proximal flange with two subequal, large spinelike processes (Fig. 149).......... Hypatopa hecate

- Ventrolateral margin of proximal flange entire, crenulate, or serrate (Figs. 151, 157, 159, 161, 165, 167, 179, 181, 185, 211, 213, 215)........................................................................................... 25

25. Margin of proximal flange entire or crenulate.............................................................. 26

- Margin of proximal flange serrate........................................................................ 30

26. Base of phallus widely bulbous basally (Figs. 152, 180, 214).................................................. 27

– Base of phallus not widely bulbous basally (Figs. 160, 166)................................................... 29

27. Vesica with cornuti (Fig. 152)................................................................. Hypatopa acus

- Vesica without cornuti (Fig. 180, 214).................................................................... 28

28. Proximal flange elongate (Fig. 213); sclerite of phallus straight basally (Fig. 214)......................... Hypatopa vox

- Proximal flange ellipsoid (Fig. 179); sclerite of phallus acutely curved basally (Fig. 180)................. Hypatopa agnae

29. Anellus 1/2 length of phallus, nearly parallelsided (Fig. 166)....................................... Hypatopa dolo

- Anellus less than 1/2 length of phallus, wider on basal 2/3, abruptly narrowed apically (Fig. 160)......... Hypatopa arxcis

30. Juxta divided mesially (Figs. 161, 211).................................................................... 31

- Juxta bandlike (Figs. 157, 167, 181, 185, 215).............................................................. 32

31. Apical process of ventral part of valva downcurved (Fig. 161); anellus broadly rounded apically (Fig. 162).... Hypatopa caedis

- Apical process of ventral part of valva upturned (Fig. 211); anellus broadly emarginate apically (Fig. 212).... Hypatopa leda

32. Anellus apically emarginate (Figs. 158, 182, 216)........................................................... 33

- Anellus apically rounded (Figs. 168, 186)................................................................. 35

33. Anellus shallowly emarginate apically, bulbous basally (Fig. 158)................................... Hypatopa hera

- Anellus deeply emarginate apically, not bulbous basally (Figs. 182, 216)......................................... 34

34. Emargination of anellus forming two apically rounded lateral lobes (Fig. 182)........................ Hypatopa phoebe

- Emargination of anellus forming two convergent, apically pointed lateral lobes (Fig. 216)............ Hypatopa tapadulcea

35. Proximal flange subrectangular; spinelike process of ventral part of valva rounded apically (Fig. 167)....... Hypatopa cyane

- Proximal flange semicircular; apical process of ventral part of valva acute (Fig. 185)..................... Hypatopa edax

36. Gnathos with a small or protuberant mesial lobe (Figs. 137, 147, 155, 173, 187)................................... 37

- Gnathos with mesial lobe absent (Figs. 141, 143, 175, 193, 197, 199, 207, 217, 227, 233)........................... 41

37. Proximal flange of valva with 1–4 spinelike processes on apicoventral region (Figs. 148, 174)........................ 38

- Proximal flange of valva without such processes (Figs. 137, 155, 187)........................................... 39

38. Proximal flange of valva elongate, densely setose (Fig. 173); sclerite of phallus broadly curved (Fig. 174).. Hypatopa cotytto

- Proximal flange of valva not elongate, sparsely setose (Fig. 147); sclerite of phallus acutely curved at 2/3 (Fig. 148)................................................................................................. Hypatopa pica

39. Margin of proximal flange of valva entire (Fig. 137); anellus with apical barb (Fig. 138)................... Hypatopa nex

- Margin of proximal flange serrate (Figs. 155, 187); anellus without apical barb (Figs. 156, 188)...................... 40

40. Proximal flange serrate (Fig. 155); apical margin of anellus narrower than base (Fig. 156)................ Hypatopa limae

- Proximal flange deeply serrate (Fig. 187); anellus truncate apically (Fig. 188)........................ Hypatopa joniella

41. Apical process of ventral part of valva diminutive (Figs. 141, 143); apex of anellus bearing a pair of recurrent lateral barblike processes (Figs. 142, 144).............................................................................. 42

- Apical process of ventral part of valva of normal length (Figs. 197, 199, 207, 217, 227, 233); anellus not bearing recurrent barblike processes (Figs. 198, 200, 208, 218, 228, 234)....................................................... 43

42. Anellus with crenulate barblike processes (Fig. 142).............................................. Hypatopa juno

- Anellus with straight barblike processes (Fig. 144)............................................... Hypatopa actes

43. Anellus mesially emarginate apically (Figs. 228, 234)........................................................ 44

- Anellus not mesially emarginate apically (Figs. 176, 194, 198, 200, 208, 218)..................................... 45

44. Anellus marginate, forming two large, divergent apicolateral processes (Fig. 228)................... Hypatopa bilobata

- Anellus marginate, forming two short apicolateral processes (Fig. 234)............................... Hypatopa musa

45. Phallus widely bulbous basally (Figs. 194, 198, 200)......................................................... 46

- Phallus not widely bulbous (Figs. 176, 208, 218)............................................................ 48

46. Apical part of sclerite of phallus slightly curved (Fig. 194).......................................... Hypatopa texla

- Apical part of sclerite of phallus recurved (Figs. 198, 200).................................................... 47

47. Basal ridge of digitate process of dorsal part of valva densely setose; base of apical process of ventral part of valva with a set- ose lobe (Fig. 199)......................................................................... Hypatopa mora

- Basal ridge of digitate process of dorsal part of valva sparsely setose; base of apical process of ventral part of valva without a setose lobe (Fig. 197).................................................................. Hypatopa tapadulcea

48. Apical process of ventral part of valva downcurved (Fig. 217)...................................... Hypatopa dicax

- Apical process of ventral part of valva not downcurved (Figs. 175, 207)......................................... 49

49. Margin of proximal flange entire (Fig. 175); sclerite of phallus shorter than phallus (Fig. 176)............ Hypatopa lucina

- Margin of proximal flange serrate (Fig. 207); sclerite of phallus as long as phallus (Fig. 208)................ Hypatopa fio

50. Eighth tergum with a darkly pigmented median longitudinal streak (Figs. 265, 270, 272–273)........................ 51

- Eighth tergum without a darkly pigmented median longitudinal streak (Figs. 261–264, 266–269, 271).................. 54

51. Ductus bursae coiled throughout most of length (Fig.265)...................................... Hypatopa tapadulcea

- Ductus bursae not coiled throughout most of length (Figs. 270, 272–273)........................................ 52

52. Posterior margin of seventh sternum emarginate (Figs. 270, 272)............................................... 53

- Posterior margin of seventh sternum straight (Fig. 273)............................................. Hypatopa vitis

53. Posterior margin of seventh sternum laterally emarginate, forming a broad, truncate, median lobe (Fig. 270).... Hypatopa rea

- Posterior margin of seventh sternum mesially and laterally emarginate, forming two rounded lobes (Fig. 272)... Hypatopa gena

54. Sex scales on membrane adjacent to Ostium bursae or on margin of seventh sternum (Figs. 261–262, 268).............. 55

- Sex scales absent (Figs. 263–264, 266–267, 269, 271)........................................................ 57

55. Sex scales on membrane adjacent to Ostium bursae (Fig. 261)..................................... Hypatopa umbra

- Sex scales on or near posterior margin of seventh sternum only (Figs. 262, 268)................................... 56

56. Sex scales on posterolateral margin of seventh sternum; antrum present (Fig. 262)..................... Hypatopa hecate

- Sex scales along entire posterior margin of seventh segment; antrum absent (Fig. 268)................... Hypatopa rabio

57. Signum present (Figs. 264, 266, 269, 271)................................................................. 59

- Signum absent (Fig. 263, 267).......................................................................... 58

58. Eighth sternum subquadrate; ductus seminalis anterior to anterior margin of seventh sternum (Fig. 267)...... Hypatopa texo

- Eighth sternum longer than wide; ductus seminalis posterior to anterior margin of seventh sternum (Fig. 263)..................................................................................................... Hypatopa plebis

59. Eighth sternum quadrate (Fig. 269)............................................................. Hypatopa sais

- Eighth sternum longer than wide (Figs. 264, 266, 271)....................................................... 60

60. Antrum present (Figs. 264, 266)......................................................................... 61

- Antrum absent (Fig. 271).................................................................... Hypatopa hora

61. Antrum wider than long; anterior 1/4 of ductus bursae spinulate (Fig. 266)............................. Hypatopa leda

- Antrum longer than wide; anterior 1/2 of ductus bursae with two rows of internal platelets (Fig. 264)...... Hypatopa phoebe