Goeldia luteipes (Keyserling, 1891)

(Figs 14; 27)

Titanoeca luteipes Keyserling, 1891: 156, pl. 4, fig. 108 (holotype male from Rio Grande (32°2’ 6”S 52°5’56”W), Rio Grande do Sul, Brazil, deposited at NHM, examined).

Calleva paupercula Simon, 1892: 239 (holotype juv ♀. from Cacheuta (33°06’S 69°07’W), Mendoza, Argentina, deposited at MNHN AR807, examined); Lehtinen 1967 (erroneously placed as syn. of G. obscura). Syn. nov.

Auximus luteipes: Petrunkevitch 1911: 106.

Amaurobius luteipes: Mello-Leitão 1919: 467, 475; Bonnet 1955: 290.

Amaurobius cothurnatus Mello-Leitão, 1943: 102, fig. 2 (holotype male from Córdoba (31°25’S 64°11’W), Argentina, deposited at MLP 15620, examined by photo); Lehtinen 1967 (Syn.).

Amaurobius thoracicus Mello-Leitão, 1945: 232 (holotype juv. from Argentina (37°11’0”S, 67°22’0”W), deposited at MLP 16345, examined). Marusik et al. 2020: 54 (sp. inquirendae). Syn. nov.

Goeldia luteipes: Lehtinen 1967: 236; Leech 1972: 103; World Spider Catalog 2023.

Additional Material Examined. VENEZUELA. Bolivar: Gran Sabana, Parupá (5°40’59”N 61°34’59”W),elev. 1500 m, 2♂ 27.VI–10.VII.1987, S. J. Peck coll.(AMNH) . BRAZIL. Amapá: Macapá (0°02’18.84”N 51°03’59.10”W), 1♂, 2–28.I.2004, R. A. Da Silva coll. (MCTP 0106580) . Paraíba: João Pessoa, Campus UFPB (7°8’22”S 34°50’43”W), 1♂, 2005–2006, P. L.F. Duarte coll. (IBSP 62904) . Distrito Federal: Brasília (15°47’38”S 47°52’58”W), 1♂, 1.XII.2000, A. J. Santos coll. (IBSP 37516) ; Área da Marinha (16°0’55”S 48°0’47”W), 1♂, 29–31.X.1999, G. G. Montingelli coll. (IBSP 26545) ; Reserva Ecológica do Instituto Brasileiro de Geografia e Estatística (15°56’41”S 47°53’07”W), 3♂, III–IV.2000, Briani coll. (IBSP 28384) . Minas Gerais: Belo Horizonte, Campus Universidade Federal de Minas Gerais (19°52’19”S 43°57’58”W), 1♂, 3.II.02, E.S.S. Álvares coll. (IBSP 73185); Estação Ecológica da Universidade Federal de Minas Gerais (19°52’S 43°58’W), 3♂, 27.III.2000, 14♂, VII.1999 – II.2001, E.S.S Álvares & E.O. Machado coll. (IBSP 27234 ‒27235, 32290); Jardim Zoológico (44°00’32”W 19°51’41”S), 1♂, 13.X.2007, A.J. Santos coll. (UFMG 1024) ; Ouro Preto, Parque Estadual do Itacolomi (20°25’45.9”S 43°28’28.7”W), 1♂, 3♀, 2–4.XI.2007, A.J. Santos et al. coll. (UFMG 1025) ; Santana do Riacho, Parque Nacional Serra do Cipó, Portaria Palácio (19°15’S 43°31’W), 1♂, 9–11.XII.2001, E.S.S. Álvares & E.O. Machado coll. (UFMG 1411) ; Santana do Riacho, Parque Nacional da Serra do Cipó (19°22’1”S 43°32’17”W), 5♂, 3♀, 2003, Equipe Biota coll. (IBSP 91607; 91609; 91611‒91612) ; Uberlândia, Reserva Ecológica do Panga (19°11’10”S 48°23’30”W), 3♂, IV.2005 – II.2006, M.F. Mineo coll. (IBSP 97338, 97340) ; Carrancas (21°29’16”S 44°38’34”W), elev. 1200 m, 1♀, X.2008, R.A. Zampaulo coll. (IBSP 124140) . Espírito Santo: Vila Velha, APA de Jacarenema (20°24’31”S 40°19’35”W), 1♂, 12.X.2005, T. Sousa coll. (IBSP 133270) . Rio de Janeiro: Macaé, Restinga de Macaé (22°22’15”S 41°47’13”W), 23.VII.1975, L. Moojer coll. (MNRJ 3216) ; Pinheiral (Currently Pinheiro) (22°30’46”S 44°00’03”W), 1♂, (MNRJ 926) ; Nova Iguaçu (22°45’37.16”S 43°26’51.82”W), 1♀, 1 juv., Blane coll. (MNRJ 929) . São Paulo: Itirapina, Estação Ecológica de Itirapina (22°15’S 47°49’W), 9♂, 1 juv., 4–9.X.2001, C. Bertin coll. (IBSP 72122–72124, 72126–72131) ; 3♂, 2001–2002, Equipe Biota coll. (IBSP 97315–97317); 10♂, 24.III– 06.XII.2000, G. Machado coll. (IBSP 91614–91611, 91622); Campinas (22°54’21”S 47°3’39”W), 1♂, 23.X.1993 (IBSP 26641) ; 1♂, 16.VIII.2006 (IBSP 79308); 1♂, 12.I.1997 (IBSP 26791); 1♂, 15.I.2002 (IBSP 35150), all collected by A.J. Santos; 1♂, X.1998, A.D. Brescovit coll. (IBSP 20129); Campus da Universidade Estadual de Campinas (22°49’3”S 47°4’11”W), 1♂, 1.II.2005, A.J. Santos coll. (IBSP 56973) ; 1♂, 18.X.2000, A.J. Santos coll. (IBSP 35131); São Carlos, Campus da Universidade Federal de São Carlos (21°58’S 47°51’W), 2♀, 1♂, VIII.2006, M. Rezende coll. (IBSP 88008‒88009) ; São Paulo, Campus da Universidade de São Paulo (23°33’44”S 46°43’39”W), 1♂, VIII.1972, Students ICB-USP coll. (IBSP 4103); 1♂, VIII.2005, G.R.S. Ruiz coll. (IBSP 56129); 1♂, G.R.S. Ruiz coll. (IBSP 59539); 1♂, 22.IX.2006, G.R.S. Ruiz coll. (IBSP 72063); 1♂, 25.IV.2000, A.D Brescovit coll. (IBSP 27291); Osasco (23°31’58”S 46°47’31”W), 1♂, 1♀, III.1991, J. Bedana coll. (IBSP 5049) ; Jaboticabal (21°15’18”S 48°19’19”W), 7♂, III– IV/2004, F.J. Cividanes coll. (IBSP 55860, 55899, 55887) ; Jundiaí, Reserva Natural Municipal Serra do Japi (23°14’0”S 46°58’0”W), 1♂, III.2007 A.D. Brescovit coll. (IBSP 78555) . Paraná: Ponta Grossa (25°05’42”S 50°09’43”W), 1♂, 2008, I.M Bortolin coll (IBSP 145404) . Santa Catarina: Rio Negrinho (26°15’14”S 49°31’04”W), 5♂, 2 juv., M.A. Nieckel coll. (IBSP 68721, I76007) . Rio Grande do Sul: Porto Alegre (30°01’58”S 51°13’48”W), 1♂, 3.IX.1983, A.D. Brescovit coll. (MCN 12664) ; Guaíba (30°06’50”S 51°19’30”W), 1♂, 4.III.1976, H. Gastal & M.H. Galileo coll. (MCN 27230) ; Bom Jesus (28°40’04”S 50°25’01”W), Fazenda Santa Cruz, 1♀, 28–30.III.1998, A.B. Bonaldo coll. (MCN 29267) , Torres (29°20’06”S 49°43’37”W), 1♂, 29.XII.1980 – 2.I.1981, P. Goloboff coll. (MACN 16121) ; Eldorado do Sul (29°59’49.7”S 51°18’28.3”W), 2♂, 25–30.XI.1995, G. Carvalho & R. Silva coll (MCTP 16528, 12560) . BOLIVIA: Cochabamba: Cochabamba (17°23’37”S 66°9’25”W), 1♀, 1940, J. Vellard coll. (MACN 16090); Monte Puncu (17°31’59”S 65°13’59”W), elev. 2700 m, 2♀, 22.X.1983, A. Roig. coll. (MACN 16024) . ARGENTINA. Tucumán: Tucumán (27°00’S 65°30’W), 6.25 Km N of Trancas, 1♀, 13.II.1951, Ross & Michaelbacher coll. (CAS 9025978) . Santiago del Estero: Moreno, Weisburd (27°18’S 62°36’W), 1♀, 26.IX.1947 (MACN 16110) . Misiones: Misiones, Santa Maria (27°54’00”S 55°23’00”W), 1♀, XI–XII.1952, J. Viana coll (MACN 16099) ; Chaco: Chaco (27°27’05”S 58°59’12”W) 1♀, 1946, G. Pinedo coll. (MACN 1735) . Córdoba: Córdoba (31°25’00”S 64°11’00”W), 2♀, (MNRJ 686) ; 1♀, J. Viana coll. (MACN 16103); 12♀, II.1952, J. Viana coll. (MACN 16033); Copina (31°33’S 64°42’W), 2♀, 2♂, XII.1983, B. R. de Fernandes coll. (MACN 16047) ; Cerro Colorado (30°05’49”S 63°55’52”W), 14.X.1961, A. de Ferraris coll. (AMNH) ; Ambúl (31°29’20”S 65°03’18”W), 2♀, 5.II.1981. A. Roig coll. (MACN 16128) ; Calamuchita (31°59’00”S 64°22’00”W), El Souce, 1♀, XII.1941, J. Viana coll. (MACN 2011) ; 1♀, XII.1941, M. J. Viana coll. (MACN 161183); Calamuchita (31°59’00”S 64°22’00”W), 4♀, 1♂, I.1955, (MACN 16031) ; 1♀, XII.1941 (MACN 16106); 2♀, XII.1941 (MACN 16104); 2♀, III.1957, (MACN 16081), all collected by J. M Viana; Cosquín (31°14’00”S 64°27’00”W), Camiño a la Mandinga, 1♂, 10.IX.1990, A. Peretti coll. (MACN 16049) ; Alta Gracia (31°40’00”S 64°26’00”W), 3er paredón, 1♂, XI.1985, M. E. Galiano & F. Miranda coll. (MACN 16108) ; La Cumbre (31°06’00”S 64°28’59”W), 1♂, 8.XI.1991, M. Ramírez coll. (MACN 16043) ; Mendoza: Mendoza, Cacheuta (32°53’00”S 68°50’00”W), 2♀, 4 juv. (MNHN 290) . San Luis: Villa Elena, Cortaderas (32°30’S 65°00’W), 1♀, 10.XI.1982, E. Maury coll. (MACN 16057) ; Villa Elena, 1♂, X.1971, J. M. Viana coll. (MACN 16093) . Buenos Aires. Tornquist, Sierra de la Ventana, Abra de la Ventana (38°04’00”S 61°58’59”W), 3♂, 2♀, 19.I.1970, M.E. Galiano coll. (MACN 16208) ; Sierra de la Ventana (38°09’S 61°48’W), Cerro Negro, 1♀, 12.IV.1974, C. Cesari coll. (MACN 16135); Tigre (34°25’00”S 58°35’00”W), 1♀, 1955, J.M. Viana coll. (MACN 16105) . URUGUAY. Canelones: Marindia (34°46’29”S 55°49’13”W), 3♂, 16.I.1997, Costa coll. (FCEN) ; Villa Argentina (34°46’00”S 55°47’00”W), 1♂, 8.II.1970, R. Capocasale coll. (CAS 9025953) .

Diagnosis. Goeldia luteipes differs from other species of the genus by the narrow, long, and triangular tegular process (Fig. 14A) and by the strongly bent tibia in the male palp (Fig. 14B–C), and by the black or dark brown color of the proximal half portion of the femora. Females of G. luteipes have a wide epigynum that covers the copulatory opening area ventrally (Fig. 14E); and copulatory ducts converging anteriorly (Fig. 14F).

Description. Male (IBSP 26545). Coloration: Carapace orange with cephalic area black; palp black; apex of coxa, trochanter, and proximal half of femora black; other leg segments and distal half of femora pale yellow except in leg IV where apex of patella to tarsus black. Opisthosoma black. Total length: 5.64. Carapace: length 2.6, width 1.84, height 1.18. Opisthosoma: length 3.04; width 1.28. Clypeus: height 0.22. Eye diameters and interdistances: AME 0.08; ALE 0.11; PME 0.95; PLE 0.08; AME–AME 0.13; AME–ALE 0.10; ALE–PLE 0.04; PLE–PME 0.19; PME–PME 0.14; AME–PME 0.11. Chelicerae paturon: length 1.41. Leg measurements: I: femur 2.1/ patella 0.72/ tibia 2.02/ metatarsus 1.94/ tarsus 0.92/ total 7.45; II: 1.68/ 0.725/ 1.64/ 2.14/ 0.93/ 7.11; III: 1.76/ 0.72/ 1.5/ 1.4/ 0.63/ 6.01; IV: 2.26/ 0.8/ 2.22/ 1.84/ 0.78/ 7.9. Leg formula: 4123. Leg spination: femur I p1ap, II p1ap; tibia II v1r-2(1r-1p), p1ap, r1ap; III v1-2 or v2sa; r1ap; p1-1ap; IV v1-1-2 or v0-1p-2, p0-1-1ap, r0-0-1ap; metatarsus I v2-1p-2(1r-1p)-2ap or v2-2(1r-1p)-2(1r-1p)- 1m, r1ap, p1ap; II v2-2(1p- 1m)-2(1r- 1m)- 1m or v2-2-2- 1m, r1ap; p1ap; d0-2-1p or d0-1p-1p; III v2-2-2- 1m; p1ap or p1-0; r1ap; d1-2-1r or d0-2-1r; IV v2-0-2-2-2- 1m, p1-2-0-1ap or p1r-1p-1r-1p-2, r1ap or r0-1r-1, d2ap; tarsus II v2sa; r0-1; III v0-2-2 or v0-1p-2, p0-1; r0-1; IV v0-2-0, r0-2-2 or r0-0-1-1, p0-1-0. Palp with patellar apophysis short and thick (Fig. 14B–C); MLT apex partially sclerotized and well developed (Fig. 14C). Pars pendula thickening subdistal (Fig. 14D). Spermatic duct bent over its on axis (Fig. 14B).

Female (IBSP 64628). Coloration: carapace brown; coxa to proximal half of the femur brown; distal half of femur and other leg segments pale brown; cephalic area, endites and chelicerae brown, slightly darker than thoracic area; opisthosoma brown. Total length: 5.25. Carapace: length 2.52, width 1.68, height 1.32. Opisthosoma: length 2.94, width 2.16. Clypeus: height 0.15. Eye diameters and interdistances:AME 0.09; ALE 0.14; PME 0.11; PLE 0.1; AME–AME 0.09; AME–ALE 0.11; ALE–PLE 0.05; PLE–PME 0.19; PME–PME 0.18; AME–PME 0.11. Paturon: 1.2 long. Leg measurements: I: femur 1.9/ patella 0.83/ tibia 1.55/ metatarsus 1.45/ tarsus 0.73/ total 6.43; II 1.76/ 0.84/ 1.47/ 1.22/ 0.78/ 6.07; III 1.48/ 0.81/ 1.0/ 0.91/ 0.63/ 4.83; IV 1.85/ 0.8/ 1.56/ 1.32/ 0.64/ 6.17. Leg formula: 1423. Leg spines: absent. Copulatory ducts close together, far from copulatory openings and converging toward middle (Fig. 14F). Extra lobe of spermathecae short (Fig. 14F)

Variation. Length (10 ♂): total: 5.20–6.63; carapace: 2.30–3.00; femur I: 1.70–2.88. (10 ♀): total 4.10–5.95; carapace 2.05–2.65; femur I 1.38–1.84.

Distribution. Venezuela, Brazil, Bolivia, Argentina, and Uruguay (Fig. 27).