Epacanthaclisis Okamoto, 1910

Epacanthaclisis Okamoto, 1910: 285 . Type species: Acanthaclisis moiwanus (Okamoto, 1905) (as Acanthaclisis moiwasana [sic] Matsumura, 1910), by original designation. Synomymized by Stange (2004: 87).

Epicanthaclisis Banks 1940: 194; Yang 1987: 212. Misspelling. Synonymized by Stange (2004: 87). Botuleon Yang, 1986: 423 . Type species: Botuleon maculosus Yang; by original designation. Synonymized by Stange (2004: 87).

Distribution. Afghanistan, Kazakhstan, Kirghizstan, Tadzhikistan, China, Japan.

Diagnosis. Bifurcation of Rs opposite or before cubital fork on wings; costal area of forewing biareolate, that of hindwing with single row of cells; 4–6 presectoral crossveins in forewing, 1–2 (usually 2) presectoral crossveins in hindwing. Female: lateral gonapophysis large, with digging setae; posterior gonapophysis smaller than anterior gonapophysis or absent. Male: pilula axillaris present; some species male with abdominal hair-pencils or tuft on IV–VI segments; genitalia without mediuncus between a pair of parameres.

Remarks. Krivokhatsky (1998) described the male genitalia of Epacanthaclisis with the mediuncus situated between the pair of parameres, but the figures of male genitalia by Krivokhatsky were not labeled. We could not find the mediuncus on these figures. So we speculate that the sclerite named mediuncus by Krivokhatsky is not homologous with the mediuncus defined by Aspöck et al. (1980). Male of some species with hair-pencils or tuft, usually situated on IV–VI abdominal segments; presumed functions of these structures could be stridulation or sex pheromone volatilization (Krivokhatsky, 1998) and may play an important role in courtship (New, 1981).