Cottus gulosus gulosus (Girard 1854), San Joaquin Riffle Sculpin, nominate subspecies

Description is the same as for the C. gulosus (Fig. 1)

Holotype: USNM 291. Same as for C. gulosus, above.

Paratype: None designated

Diagnosis: Distinguished from other members of the C. gulosus complex as a distinct lineage as determined by genomic studies (Baumsteiger et al. 2014, this study) and by its limited geographic distribution (Fig. 3). See species description for distinguishing it from co-occurring C. asper .

Distribution. These California endemic sculpins are found in cold-water riffles, in headwaters or below dams with cold water releases into rivers in the southern Sierra Nevada on the eastern side of the Central Valley, except for the American River (which flows directly into the Sacramento River), the rivers that contain now-isolated populations all flow into the southern Central Valley, especially the San Joaquin River. Rivers with populations include the Mokelumne, Stanislaus, Tuolumne, Merced, Fresno, San Joaquin, Kings, and Kern rivers.

Etymology. See C. gulosus