Eupines (Byraxis) glabrata (Broun, 1886)

Fig. 19; map Fig. 51

Bryaxis glabrata Broun, 1886: 830 .

Eupines (Byraxis) glabrata: combination and subgeneric assignment by Raffray 1904: 207. Raffray 1911: 82, 1924: 157; Hudson 1923: 364, 1934: 182; Kuschel 1990: 49; Nomura and Leschen 2006: 254; Maddison 2010: 417; Shen and Leschen 2019: 265.

Type locality: [AK] Woodhill, near the Kaipara Railway.

Type depository: NZAC HT ♂ .

Diagnosis: Head with vertexal foveae punctiform; antennae 10-segmented with 4-segmented antennal club; A4, A5, A6, A7, A8 and A10 unmodified, A6 and A8 slightly asymmetrical, A9 largest, with a deep groove at one side, forming laterally twisted lamina at apex, A10 conical and much shorter than A9. Abdominal ventrite 2 with pair of small discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of short and acute median struts, about one-tenth of ventral length. Protrochanters with long and acute ventral spine, about same to protrochanteral width; protibiae and mesotibiae unmodified.

Redescription: Body length 1.33–1.44 mm; body colour reddish brown to dark brown with lighter coloured tibiae and tarsi. Head wider than long; frons not narrowed at end, about three-fourths width of head at eye level; eyes well developed, with about 31 facets; vertexal foveae punctiform; antennal tubercles moderately developed; postantennal notches broadly extended and meet at middle of vertex; postantennal cavities medium-sized and about two-thirds length of antennal tubercles in dorsal view; antennae 10-segmented with 4-segmented antennal club; A4, A5, A6, A7, A8 and A10 unmodified, A6 and A8 slightly asymmetrical, A9 largest, with a deep groove at one side, forming laterally twisted lamina at apex, A10 conical and much shorter than A9. Pronotum as long as wide, slightly narrower than head at eye level; lacking lateral antebasal foveae. Elytra longer than wide, widest at about apical three-fourths; lacking humeral calli; sutural stria barely visible. Abdomen with tergite 1 about 1.2 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with strongly curved small apical setae; ventrite 6 with pair of short and acute median struts, about one-tenth of ventral length. Legs with protrochanters modified with long and acute ventral spine, about same to protrochanteral width; protibiae and mesotibiae unmodi- fied. Aedeagus about 0.28 mm long, with parameres long and clavate at apex, curved inwards; median lobe with four short and stout sclerites originating from middle in dorsal view, two outside ones curved outwards, two middle ones strongly curved inwards.

Comments: This species is widely distributed in the North Island and is also reported in Marlborough Sounds. It can be distinguished from congeners by its A8 conspicuously transerverse and enlarged (Fig. 19C).

Distribution: North Island. ND, AK, WO, CL, BP, GB, TK, HB, WN, WA. South Island. SD.

Additional material examined: 90 ♂♂, ND: 1, E. Pandora Rd., Te Paki Coastal Park, 07/Feb/1975, A.K. Walker; 2, Mokohinau Is., Fanal I., 01/Mar/1978, G. Kuschel, sifted litter; 6, Poor Knights Is., Tawhiti Rahi, 10/ Sep/1980, J.C. Watt litter; AK: 6, Lynfield, 27/Jan/1977, G. Kuschel, litter; 1, Noises Is., Motuhoropapa I. South Trig, 15/Jan/1978, L.L. Deitz, litter near pit trap; 1, Bethells Matuku Res., 26/Jul–20/Aug/1983, G. Hall and P.A. Maddison, pit traps; 3, Coll. A.E. Brookes, Sep/1914, Titirangi Waitakere Rg., Auckland, 1476; 10, Clevedon R.A. Cumber Coll., 09/Jul/1958; 4, AMNZ, Lynfield, 27/Jan/1977, G. Kuschel; CL: 1, Little Barrier I., Te Maraeroa South side, 07/Apr/1984, C.T. Duval, sifted litter; 2, AMNZ, Cuvier I., West Ridge Track, 160 m, 12/xi/1999, J.W. Early, S.E. Thorpe; WO: 1, Waitomo, 24/May/1983, G. Kuschel, decayed wood and litter; 1, Mt. Pirongia SF., Rangitukia stream, 13/Jun/2001, R. Leschen, leaf litter, 37°58'S 175°07'E; BP: 2, Lottin Pt Rd., Waenga Bush, 27/ Jan/1993, R.C. Henderson, litter; 1, Hicks Bay, 14/Mar–29/Apr/1993, J.S. Dugdale, litter; 9, Te Koau Twin Puriri, 31/Jan/1993, J.S. Dugdale litter; 1, Waiotuma Stm., 65 m, 28/Jan/1998, J.S. Dugdale, litter; 2, LUNZ, Te Koau, Twin Puiris, 15/iii/1993, R.M. Gilbert, J.W.M. Marris; GB: 1, East Cape, Rangiata, 17/Mar–30/Apr, G. Hall, pit traps; 2, Whakaroa Scenic Reserve, 22/Nov/1993 – 11/Jan/1994, G. Hall, pit traps; 2, Kakanui, 350 m, 01/Feb/1993, J.S. Dugdale, litter; 2, Taikawakawa, 21/Sep/1992, G. Hall & R.C. Henderson, sifted litter, 92/58; 1, LUNZ, Kakanui, 260 m 16/iii/1993, J.W.M. Marris; TK: 1, White Cliffs Walkway, 5 km S. Tongaporutu, 07/Dec/1982, J.S. Dugdale, litter 82/137; HB: 13, Motuokura (Bare I.), 07/Feb–20/Mar/1995, G. Hall, pit traps; RI: 4, Coll. J. Ford., 28/9/1917, Taihape main Trunk, N. Island, A.E. Brookes Collection; WA: 1, W Shore Res., L Wairarapa, 09/Oct/1993; WN: 6, JNPC, Nikau Reserve, Paraparaumu, 23/Jul/95, J.T. Nunn collection; SD: 1, LUNZ, Titi Island, 17/xi/1994, J.W.M. Marris, litter ex broadleaf forest and seabird burrow; 4, JNPC, Titi Island, 03/April/1990, J.I. Townsend. Unknown locality: 2, 1476, T. Broun Collection, A.E. Brookes Collection. Associated females (57 ♀♀): ND: 15, Mokohinau Is., Fanal I., 01/Mar/1978, G. Kuschel, sifted litter; AK: 17, Lynfield, 27/Jan/1977, G. Kuschel, litter; 4, Coll. A.E. Brookes, Sep/1914, Titirangi, Waitakere Rg., Auckland, 1476; 1, Bethells Matuku Res., 26/Jul–20/Aug/1983, G. Hall and P.A. Maddison, pit trap; 1, Riverhead, Forest Reserve, 30/Aug/1982, P.A. Maddison, litter; GB: 11, Taikawakawa, 21/Sep/1992, G. Hall & R.C. Henderson, sifted litter, 92/58; BP: 1, LUNZ, Te Koau, Twin Puiris, 15/iii/1993, R.M. Gilbert, J.W.M. Marris; HB: 2, Motu-okura (Bare I.), 07/Feb–20/Mar/1995, G. Hall, pit traps; SD: 5, LUNZ, Titi Island, 17/xi/1994, J.W.M. Marris, litter ex. broadleaf forest and seabird burrow.