39. Polistes (Gyrostoma) tenebricosus Lepeletier, 1836

Polistes tenebricosa Lepeletier 1836: 529 . Type locality: Java, Indonesia.

Polistes sulcatus Smith, 1852a: 38 . Type data: Female, male BMNH. Type locality: Ningpo, China.

Polistes hoplitus de Saussure, 1853: 55 . Type data: Female, MHNG. Type locality: Indes Orientales.

Polistes hoplites de Saussure, 1854: 255 .

Polistes rugifrons Cameron, 1900: 412 .

Polistes varicornis Cameron, 1905: 67 .

Polistes javanicus Cameron, 1905: 68 .

Polistes tenebricosus var. leopoldi Bequaert, 1934: 9 .

Polistes tenebricosus var. sibuyanensis Bequaert, 1940: 266 .

Distribution: Bhutan: Trashigang, Mongar, Wangduephodrang, Chhukha, Pema Gatshel, Zhemgang, Lhuentse (Dorji et al. 2016; Dorji, Klein, et al. 2017a). Elsewhere: China; India; Nepal; Vietnam; Myanmar; Indonesia; Philippines (Carpenter 1996; Das & Gupta 1989; Dorji et al. 2016; Dorji, Klein, et al. 2017a; Gawas et al. 2020; Kumar 2010; Kumar & Sharma 2014a, 2015b; Nguyen & Carpenter 2019; Pham & Li 2015; Sora et al. 2019; Tan, van Achterberg, Duan, et al. 2014; Tenzin & Katel 2019; Yamane & Yamane 1979).