TRIBE LACCOCORINI POPOV, 1970

FIGS 3B, 5C, D, 6, 10E, 11E, 14B, D, 15A, 20

Diagnosis: A unifying feature for all members of Laccocorini except Aneurocoris is that the anteroventral seta row on the metafemur gradually extends toward the middle of the ventral surface basally (Fig. 5d), or if there is an inflection in the direction, it occurs in the distal half of the femur. The spinose setae of the posteroventral row on the mesofemur are not tightly packed and are separated from each other at least by the width of a seta (Fig. 5c) and number approximately 40–60. The labrum has a transverse sulcus at the base (Fig. 6), although the sulcus appears to have been lost in Temnocoris .

Comments: Because Laccocoris spurcus is the type species for the subfamily, this clade is here recognized as tribe Laccocorini . This tribe is endemic to Africa, Madagascar and the Middle East. Generic level relationships among Aneurocoris, Laccocoris, Temnocoris and the new genus Tsingala are strongly supported (bs = 100). The clade containing Aneurocoris and Laccocoris from mainland Africa have a unique fracture (Fig. 10e) across the hemelytra from the distal ends of the embolia through the tip of the claval commissure (Mbogho & Sites, 2013). This fracture is present in both sexes of Aneurocoris insolitus Montandon, 1897b and mostly only females of Laccocoris, although it is also present in some males. The Middle Eastern Heleocoris minisculus Walker, 1870 also bears this fracture and is here transferred to Laccocoris as L. minisculus (Walker, 1870) comb. nov., but the fracture is absent in the clade of Temnocoris + Tsingala from Madagascar.

Included taxa: Aneurocoris (known by only two species in Lake Tanganyika) and Laccocoris are from mainland Africa, and Temnocoris and Tsingala are endemic to Madagascar.