TRIBE IDIOCARINI TRIB. NOV.

FIG. 23

Z o o b a n k r e g i s t r a t i o n: u r n: l s i d: z o o b a n k. org:act: 3F13F62A-0CCB-4452-9224-7CE361467AF7

Diagnosis: Despite the diversity of genera in this tribe, a synapomorphy that unites them is the shape of the left paramere of the males (Fig. 23c). Specifically, they tend to be relatively broad and lack the pronounced lobe of the left side that is present in the other tribes of Naucorinae (Figs 22c, 24e), thus rendering the structure as appearing less modified, whereas this is probably the derived character state given its phylogenetic position in the dendrogram and that the lobe is present in the other tribes. Species of Sagocoris may have secondarily acquired an incipient lobe unrelated to the pronounced lobe of the other tribes.

Comments: The seven New Guinean endemic genera of the former tribes Sagocorini and Tanycricini are here transferred from Cheirochelinae to Naucorinae . They were recovered with an Australian species of Naucoris at the root. For this strongly supported clade (pp = 1.0, bs = 100), I propose the new tribe Idiocarini and Australonaucoris gen. nov. described below, in which to transfer Naucoris congrex . In addition to N. congrex, five other species of Naucoris are known from Australia: Naucoris australicus Stål, 1876, Naucoris magela Lansbury, 1991, Naucoris scutellaris (= Naucoris rhizomatus J. Polhemus, 1984; some authorities placed this species in Thurselinus Distant, 1904), Naucoris subaureus Lansbury, 1985 and Naucoris subopacus Montandon, 1913 .

Included taxa: This tribe contains seven genera endemic to New Guinea, including Aptinocoris Montandon, 1897d, Cavocoris, Idiocarus, Nesocricos, Sagocoris, Tanycricos and Warisia La Rivers, 1971 . Also included is Australonaucoris from Australia. Three of the other species of Naucoris from Australia also are transferred to Australonaucoris (see description of the genus below).